After Widmer, Lötters & Jungfer, 2000, with the Recent species from Frost, 2000-2002 [“AMNH Amphibian species of the World”]
<==o Dendrobatidae (dart-poison frogs; nuolimyrkkysammakot) |--o Allobates Zimmermann & Zimmerman, 1988 [Epipedobates femoralis -group] | |-- A. boulengeri (Barbour, 1909) Myers, 1987 [taxonomic history] | |-- A. espinosai (Funkhouser, 1956) Myers, 1987 [taxonomic history] | |-- A. femoralis (Boulenger, 1884) Myers, 1987 [taxonomic history] | |-- A. myersi (Pyburn, 1981) Myers, 1987 [taxonomic history] | |-- A. tricolor (Boulenger, 1899) Myers, 1987, Rivero & Almendáriz [taxonomic history] | `-- A. zaparo (Silverstone, 1976) Myers, 1987 [taxonomic history] |?- Aromobates nocturnus (Meyers, Paolillo & Daly, 1991) |?- Colostethus Cope, 1866 [taxomonic history] ()? [99] |?-o Mannophryne La Marca, 1992 [taxonomic history] ()? [12] | |-- M. caquetio Mijares-Urrutia and Arend, 1999 | |-- M. collaris (Boulenger, 1912) La Marca, 1992 [taxonomic history] | |-- M. cordilleriana La Marca, 1995 | |-- M. herminae (Boettger, 1893) La Marca, 1995 [taxonomic history] | |-- M. lamarcai Mijares-Urrutia and Arend, 1999 | |-- M. larandina (Yústiz, 1991) Mijares-Urrutia and Arend, 1999 [taxonomic history] | |-- M. neblina (Test, 1956) La Marca, 1992 [taxonomic history] | |-- M. oblitterata (Rivero, 1986) La Marca, 1992 [taxonomic history] | |-- M. olmonae (Hardy, 1983) La Marca, 1992 [taxonomic history] | |-- M. riveroi (Donoso-Barros, 1965) La Marca, 1992 [taxonomic history] | |-- M. trinitatis (Garman, 1888) La Marca, 1995 [taxonomic history] | `-- M. yustizi (La Marca, 1989) La Marca, 1992 [taxonomic history] |?-o Minyobates Myers, 1987 [= Dendrobates Wagler, 1830] | |-- M. abditus (Myers & Daly, 1976) Meyers, 1987 [Dendrobates abditus Myers & Daly, 1976, Dendrobates abditus Myers & Daly; Jungfer, Lötters, Jörgens, 2000] | |-- M. altobueyensis (Silverstone, 1975) Meyers, 1987 [Dendrobates altobueyensis Silverstone, 1975; Dendrobates altobueyensis Silverstone; Jungfer, Lötters, Jörgens, 2000] | |-- M. bombetes (Myers & Daly, 1980) Meyers, 1987 [Dendrobates bombetes Myers & Daly, 1980; Dendrobates bombetes Myers & Daly; Jungfer, Lötters, Jörgens, 2000} ] | |-- M. fulguritus (Silverstone, 1975) not Meyers, 1987 [Dendrobates fulguritus Silverstone, 1975; Dendrobates fulguritus Silverstone; Jungfer, Lötters, Jörgens, 2000] | |-- M. minutus (Shreve, 1935) Meyers, 1987 [Dendrobates minutus Shreve, 1935; Dendrobates minutus Shreve; Jungfer, Lötters, Jörgens, 2000; Dendrobates minutus minutus Shreve, 1935; Dendrobates shrevei Dunn, 1940] | |-- M. opisthomelas (Boulenger, 1899) Meyers, 1987 [Dendrobates opisthomelas Boulenger, 1899; Dendrobates opisthomelas Boulenger; Jungfer, Lötters, Jörgens, 2000] | |-- M. steyermarki (Rivero, 1971) Meyers, 1987 [Prostherapis steyermarki Rivero, 1971, Dendrobates steyermarki (Rivero); Jungfer, Lötters, Jörgens, 2000] | |-- M. viridis (Myers & Daly, 1976) Meyers, 1987 [Dendrobates viridis Myers & Daly, 1976; Dendrobates viridis Myers & Daly; Jungfer, Lötters, Jörgens, 2000] | `-- M. virolensis Ruiz-Carranza & Ramírez-Pinilla, 1992 [D. virolinensis (Ruiz-Carranza & Ramírez-Pinilla) Jungfer, Lötters, Jörgens, 2000] |?-o Nephelobates La Marca, 1992 ()? [9] | |-- N. alboguttatus (Boulenger, 1903) La Marca, 1994 [taxonomic history] | |-- N. duranti (Péfaur, 1985) La Marca, 1997 [taxonomic history] | |-- N. haydeeae (Rivero, 1978) La Marca, 1994 [taxonomic history] | |-- N. leopardalis (Rivero, 1978) La Marca, 1991[taxonomic history] | |-- N. mayorgai (Rivero, 1980) La Marca, 1994 [taxonomic history] | |-- N. meridensis (Dole & Durant, 1972) La Marca, 1994 [taxonomic history] | |-- N. molinarii (La Marca, 1985) La Marca, 1994 [taxonomic history] | |-- N. orostoma (Rivero, 1978) La Marca, 1994 [taxonomic history] | `-- N. serranus (Péfaur, 1985) La Marca, 1997 [taxonomic history] |-- Epipedobates Myers, 1987 ()? [30] `--+-- Dendrobates Wagler, 1830 ()? [26] `--o Phyllobates Dumeril & Bibron, 1841 ()? [5] |--+-- P. lugubris (Schmidt, 1857) Dunn, 1924 [taxonomic history] | `-- P. vittatus (Cope, 1893) Silverstone, 1975 [taxonomic history] `--+-- P. aurotaenia (Boulenger, 1913) Dunn, 1957 [taxonomic history] `--+-- P. bicolor (Duméril & Bibron, 1841) [taxonomic history] `-- P. terribilis Myers, Daly & Malkin, 1978
Reference(s):
- Frost, D. R., 2000-2003: Amphibian species of the World: An Online Reference.
–iNet: American Museum of Natural History, Department of Herpetology: http://research.amnh.org/herpetology/amphibia/index.html - Grant, T., Frost, D. R., Caldwell, J. P., Gagliardo, R., Haddad, C. F. B., Kok, P. J. R., Means, D. B., Noonan, B. P., Schargel, W. E. & Wheeler, W. C., 2006: Phylogenetic systematics of dart-poison frogs and their relatives (Amphibia: Athesphatanura: Dendrobatidae).
–Bulletin of the American Museum of Natural History: Vol. #299, pp. 1-262 - Widmer, A., Lötters, S. & Jungfer, K.-H., 2000: A molecular phylogenetic analysis of the neotropical dart-poison frog genus Phyllobates (Amphibia: Dentrobatidae).
–Naturwissenschaften: Vol. 87, #11, pp. 559-562