After Grant et al., 2006, with the Recent species from Frost, 2000-2002 [“AMNH Amphibian species of the World”]
<==o Aromobatidae Grant et al., 2006 |--o Anomaloglossinae Grant et al., 2006 | |--o Rheobates Grant et al., 2006 | | |-- R. palmatus (Werner, 1899) [Phyllobates palmatus Werner, 1899] | | `-- R. pseudopalmatus (Rivero & Serna, 2000 “1995”) | `--o Anomaloglossus Grant et al., 2006 [Colostethus Cope, 1866 (in partim)] | |-- A- astralogaster Myers, Ibáñez D., Grant & Jaramillo, 2012 | |-- A. atopoglossus (Grant, Humphrey & Myers, 1997) | |-- A. ayarzaguenai (La Marca, 1998 “1996” | |-- A. baeobatrachus (Boistel & Massary, 1999) | |-- A. beebei (Noble, 1923) | |-- A. “chocoensis” auctorum (non Boulenger, 1912) | |-- A. degranvillei (Lescure, 1975) | |-- A. guanayensis (La Marca, 1998 “1996” | |-- A. isthminus Myers, Ibáñez D., Grant & Jaramillo, 2012 [Colostethus chocoensis (Boulenger) Myers, 1991 (in partim)] | |-- A. lacrimosus (Myers, 1991) | |-- A. murisipanensis (La Marca, 1998 “1996”) | |-- A. parimae (La Marca, 1998 “1996”) | |-- A. parkerae (Meinhardt & Parmelee, 1996) | |-- A. praderioi (La Marca, 1998 “1996”) | |-- A. roraima (La Marca, 1998 “1996”) | |-- A. shrevei (Rivero, 1961) | |-- A. stepheni (Martins, 1989) | |-- A. tamacuarensis (Myers & Donnelly, 1997) | |-- A. tepuyensis (La Marca, 1998 “1996”) | |-- A. triunfo (Barrio-Amorós, Fuentes & Rivas, 2004) | `-- A. wothuja (Barrio-Amorós, Fuentes & Rivas, 2004) `--+--o Allobatinae Grant et al., 2006 | `--o Allobates Zimmermann & Zimmerman, 1988 [Epipedobates femoralis -group] | |-- A. boulengeri (Barbour, 1909) Myers, 1987 [taxonomic history] | |-- A. espinosai (Funkhouser, 1956) Myers, 1987 [taxonomic history] | |-- A. femoralis (Boulenger, 1884) Myers, 1987 [taxonomic history] | |-- A. myersi (Pyburn, 1981) Myers, 1987 [taxonomic history] | |-- A. tricolor (Boulenger, 1899) Myers, 1987, Rivero & Almendáriz [taxonomic history] | `-- A. zaparo (Silverstone, 1976) Myers, 1987 [taxonomic history] `--o Aromobatinae Grant et al., 2006 |-- Aromobates nocturnus Meyers, Paolillo & Daly, 1991 `--o Mannophryne La Marca, 1992 [taxonomic history] ()? [12] |-- M. caquetio Mijares-Urrutia and Arend, 1999 |-- M. collaris (Boulenger, 1912) La Marca, 1992 [taxonomic history] |-- M. cordilleriana La Marca, 1995 |-- M. herminae (Boettger, 1893) La Marca, 1995 [taxonomic history] |-- M. lamarcai Mijares-Urrutia and Arend, 1999 |-- M. larandina (Yústiz, 1991) Mijares-Urrutia and Arend, 1999 [taxonomic history] |-- M. neblina (Test, 1956) La Marca, 1992 [taxonomic history] |-- M. oblitterata (Rivero, 1986) La Marca, 1992 [taxonomic history] |-- M. olmonae (Hardy, 1983) La Marca, 1992 [taxonomic history] |-- M. riveroi (Donoso-Barros, 1965) La Marca, 1992 [taxonomic history] |-- M. trinitatis (Garman, 1888) La Marca, 1995 [taxonomic history] `-- M. yustizi (La Marca, 1989) La Marca, 1992 [taxonomic history]
Reference(s):
- Frost, D. R., 2000-2003: Amphibian species of the World: An Online Reference.
–iNet: American Museum of Natural History, Department of Herpetology: http://research.amnh.org/herpetology/amphibia/index.html - Grant, T., Frost, D. R., Caldwell, J. P., Gagliardo, R., Haddad, C. F. B., Kok, P. J. R., Means, D. B., Noonan, B. P., Schargel, W. E. & Wheeler, W. C., 2006: Phylogenetic systematics of dart-poison frogs and their relatives (Amphibia: Athesphatanura: Dendrobatidae).
–Bulletin of the American Museum of Natural History: Vol. #299, pp. 1-262 - Myers, C. W., Ibáñez D., R., Grant, T. & Jaramillo, C. A., 2012: Discovery of the Frog Genus Anomaloglossus in Panama, with descriptions of two new species from the Chagres Highlands (Dendrobatoidea: Aromobatidae).
–American Museum Novitates: Vol. #3763, pp. 1-19 [doi: 10.1206/3763.2]