After Yates & Warren, 2000, Steyer, 2002, Damiani, 2004, and Schoch, 2006
<==o †Trematosauroidea Watson, 1919 sensu Yates & Warren, 2000, and Damiani, 2001 |?- †Benthosphenus lozovskii Shishkin [& Lozovsky], 1979 |?-o †Thoosuchidae [paraphyletic Benthosuchidae: Thoosuchinae] | |?-o †Prothoosuchus Getmanov, 1989 | | |-- †P. blomi Getmanov, 1989 | | `-- †P. samariensis Getmanov, 1989 | `--o †Thoosuchus Efremov, 1940 | |-- †T. yakovlevi (Ryabinin, 1927) | |-- †T. tardus Getmanov, 1989 | `-- †T. tuberculatus Getmanov, 1989 `--o †Trematosauridae Watson, 1919 |--o †Trematosaurinae Watson, 1919 | |?- †Tirraturhinus smisseni Nield, Damiani & Warren, 2006 | |--+--+-- †Tertrema acuta Wiman, 1915 | | | `?-+-- †Tertremoides ambilobensis | | | `-- †Trematolestes hagdorni Schoch., 2006 [“Labyrinthodont” Meyer & Plieninger, 1844; Labyrinthodon sp. Fraas, 1889; Plagiosuchus pustuliferus Huene, 1922; Trematosauridae n. g. n. sp. Schoch & Werneburg, 1998; Trematosauridae n. g. n. sp. Schoch & Wild, 1999; Bukobaja cf. B. enigmatica Schoch, 1999; Trematosauridae n. g. n. sp. Schoch & Milner, 2000] | | `--+-- †Lyrocephaliscus euri Wiman, 1914 [Lyrocephalus Säve-Söderbergh, 1935] | | `--+-- †Platystega depressa Wiman, 1915 | | |?- †Microposaurus casei Haughton, 1925 [Rhytidosteus capensis (Owen, 1884) Romer, 1947; Rhytidosteus capensis (Owen) Huene, 1956; Rhytidosteus capensis (Owen) Kitching, 1957; Rhytidosteus capensis (Owen, 1884) Shishkin, 1964] | | `?- †Inflectosaurus amplus Shishkin, 1960 | `--+-- †Luzocephalus blomi Shishkin, 1980 [Lyzocephalidae Shishkin, 1980] | `--+--o †Trematosaurus Burmeister, 1849 [Trematosaurus von Braun, 1842; Hellrung, 1987); nomen nudum (Schoch & Milner, 2000))] | | |-- †T. brauni Burmeister, 1849 (non Wepfer, 1922) [Trematosaurus fuchsi Seidlitz, 1920] | | `-- †T. madagascariensis Lehman, 1966 [non T. madagascarensis Hewison, 1996; Tertremoides ambilobensis Lehman, 1979] | `--o †Trematosuchus [Thoosuchus] | |-- †T. sobeyi Haughton, 1915 | `-- †T. yakovlevi Riabinin, 1927 [Thoosuchus yakovlevi] `--o †Lonchorhynchinae Säve-Söderbergh, 1935 |--o †Aphaneramma Säve-Söderbergh, 1935 | |-- †A. ocella (Meyer) Huen, 1920 | `-- †A. rostratum [‘Helobatrachus’ Hammer, 1987; ‘Lonchorhynchus obergi’ Wiman, 1910; ‘Trematosaurus kannemeyeri’ Broom, 1909] `--+-- †Erythrobatrachus noonkanbahensis Cosgriff & Garbutt, 1972; Welles, 1993 [†Erythrobatrachus hoonkanbahensis Hellrung, 1987 (lapsus calami?)] |-- †Gosgriffius campi Welles, 1993 `--+-- †Wantzosaurus elongatus Lehman, 1961 [Wantzosaurus sp. Lehman, 1955; Wantzosaurus elongatus Lehman, 1961; ‘stégocéphale néoténique’ Lehman, 1963; Ifasaurus elongatus Lehman, 1966; trematosaurian incertae sedis Warren & Black, 1985] `-- †Stoschiosaurus nielseni Säve-Söderbergh, 1935
Trematosauroidea incertae sedis:
†Angusaurus tsylmensis Novikov, 1990 †Indolyrocephalus huxleyi (Lydekker, 1882) Tripathi, 1969 [Gonioglyptus huxleyi Lydekker, 1882] †Gonioglyptus kokeni Huene, 1920 †Panchetosaurus longirostris (Huxley, 1865) Tripathi, 1969 [Gonioglyptus longirostris Huxley, 1865] †Hyperokynodon kauperinus (Plieninger, 1852) [Hyperokynodon kauperinum (Plieninger); Hammer, 1987; Warren & Black, 1985] †Icanosaurus rectifrons Rusconi, 1951 †Luzocephalus johannsoni (Säve-Söderbergh, 1935) †Luzocephalus kochi (Säve-Söderbergh, 1935) †Trematotegmen ostchevi Getmanov, 1982
Reference(s):
- Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Carroll, R. L., 1988: Appendix. 594-648
in Carroll, R. L. 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698. - Damiani, R. J., 2004: Cranial anatomy and relationships of Microposaurus casei, a temnospondyl from the Middle Triassic of South Africa.
–Journal of Vertebrate Paleontology: Vol. 24, #3, pp. 533-541 - Nield, C. M, Damiani, R., & Warren, A., 2006, A short-snouted trematosauroid (Tetrapoda, Temnospondyli) from the Early Triassic of Australia: the oldest known Trematosaurinae.
–Alcheringa: Vol. 30, pp. 263-271 - Schoch, R. R., 2006: A complete trematosaurid amphibian from the Middle Triassic of Germany.
–Journal of Vertebrate Paleontology: Vol. 26, #1, pp. 29-43 - Steyer, J.-S., 2002: The first articulated trematosaur ‘amphibian’ from the Lower Triassic of Madagascar: implications for the phylogeny of the group.
–Palaeontology: Vol. 45, #4, pp. 771-793 - Yates, A. M. & Warren, A. A., 2000: The phylogeny of the "higher" temnospondyls (Vertebrata: Choanata) and its implications for the monophyly and origins of Stereospondyla.
–Zoological Journal of the Linnean Society: Vol. 128, pp. 77-121