After Schoch, 2000
<==o †Capitosauridae Watson, 1919 [basal Mastodontosauroidea sensu Damiani, 2001, and Steyer, 2003] |-- †Parotosuchus madagascariensis Lehman, 1961 `--+--o †Parotosuchus Otschev & Shishkin, 1968 [taxonomic history] | |-- †P. rajareddyi Roy-Chowdhury, 1970 [Eryosuchus rajareddyi (Roy-Chowdhury, 1970) Damiani, 2001] | |-- †P. crookshanki Mukherjee & Sengupta, 1998 [Paracyclotosaurus crookshanki (Mukherjee & Sengupta, 1998) Damiani, 2001] | |-- †P. aliciae Warren & Hutchinson, 1988 [Watsonisuchus aliciae (Warren & Hutchinson, 1988) Damiani, 2001] | |-- †P. rewanensis Warren, 1980 [Watsonisuchus rewanensis (Warren, 1980) Damiani, 2001] | |-- †P. haughtoni (Broili & Schröder, 1937) Romer, 1947 [taxonomic history] | |-- †P. helgolandicus (Schröder, 1913) Chernin, 1978 [taxonomic history] | |-- †P. komiensis Novikov, 1986 | |-- †P. nasutus (Meyer, 1858) Chernin, 1978 [taxonomic history] | |-- †P. oremburgensis (Konzhukova, 1965) Ochev & Shishkin, 1968 [taxonomic history] | |-- †P. orientalis (Otschev [Ochev], 1966) Ochev & Shishkin, 1968 [taxonomic history] | |-- †P. panteleevi (Otschev [Ochev], 1966) Ochev & Shishkin, 1968 [taxonomic history] | |-- †P. sequester Lozovsky & Shishkin, 1974 [taxonomic history] | |-- †P. speleus Shishkin & Sulej, 2009 | `-- †P. ptaszynskii Sulej & Nied´zwiedzki, 2013 `--+--o †“Mastodontosaurinae” | |--o †Mastodontosaurus Jaeger, 1828 [Salamandroides Jaeger, 1828; Batrachosaurus Fitzinger, 1840; Labyrinthodon Owen, 1842a; Heptasaurus Säve-Söderbergh, 1935] | | |?- †Mastodonsaurus vaslenensis Meyer, 1855 [Mastodonsauridae indet. sensu Damiani, 2001] | | |-- †M. cappelensis Wepfer, 1923b [Heptasaurus cappelensis (Wepfer, 1923b) Säve-Söderbergh, 1935; Mastodonsaurus ingens Trusheim, 1937] | | |-- †M. torvus Konzhukova, 1955 [M. tantus Krasilnikov & Vyushkov, 1947 nomen nudum; M. toevus Konzhukova, 1955 lapsus calami; M. maximus Ochev, 1958b] | | `-- †M. keuperinus | `--o †Eryosuchus Otschev, 1966 | |-- †E. tverdochlebovi Otschev, 1966 | |-- †E. garjainovi Otschev, 1966 | `-- †E. antiquus Otschev, 1966 `--+--+-- †Parotosuchus pronus Howie, 1970 [Eryosuchus pronus (Howie, 1970) Damiani, 2001] | `-- †Wellesaurus peabodyi (Welles & Cosgriff, 1965) Lehman, 1971 [Parotosaurus peabodyi (Welles & Cosgriff, 1965] `--+--+-- †Kupferzellia | `--o †Cyclotosaurus Fraas, 1889 [Capitosaurus Münster, 1844; Mastodonsaurus Jaeger, 1850; Paracyclotosaurus Watson, 1958] | |-- †C. ebrachensis Kuhn, 1932 | |-- †C. hemprichi Kuhn, 1942 [Paracyclotosaurus hemprichi (Kuhn, 1942) Watson, 1958] | |-- †C. papilio Wepfer, 1923a | |-- †C. robustus (Quenstedt, 1850) Fraas, 1889 [Capitosaurus robustus Mayer, 1844 nomen vanum; Mastodonsaurus robustus Quenstedt, 1850, Mastodonsaurus cyclotis Quenstedt, 1850] | |-- †C. carinidens | `-- †C. posthumus Fraas, 1913 [Paracyclotosaurus posthumus (Fraas, 1913) Shishkin, 1960; incl. C. mordax Fraas, 1913] |-- †Quasicyclotosaurus `--o †Eocyclotosaurus Ortlam, 1970 [Stenotosaurus Romer; Parotosaurus Jaekel; Heylerosaurus Shishkin, 1980] [are heylerosaurs (Yates & Warren, 2000)] |-- †E. woschmidti Ortlam, 1970 [Stenotosaurus woschmidti (Ortlam, 1970) Heyler, 1976] `-- †E. lehmani (Heyler, 1969) Kamphausen & Morales, 1981 [Stenotosaurus lehmani Heyler, 1969; Parotosaurus lehmani (Heyler, 1969) Carroll & Winer, 1977; Heylerosaurus lehmani (Heyler, 1969) Shishkin, 1980]
Capitosauridae not treated by Schoch, 2000:
†Bukobaja enigmatica (Otschev, 1966)
†Calamops paludosus
†Capitosaurus arenaceus Münster, 1836
†Capitosaurus polaris Wiman, 1915
†Cyclotosaurus spitzbergensis Wiman, 1915
†Kestrosaurus dreyeri Haughton, 1925
†Paracyclotosaurus davidi Watson, 1958
†Promastodonsaurus bellmanni
†Sassenisaurus
†Tatrasaurus Marya´nska & Shishkin, 1996
†T. kulczyckii Marya´nska & Shishkin, 1996
†T. wildi (Schoch, 1997) Damiani, 2001
†Volgasaurus
†Volgasuchus
†Watsonisuchus Ochev, 1966
†W. magnus (Watson, 1962) Ochev, 1966 [Wetlungasaurus magnus Watson, 1962]
†Xenotosaurus africanus (Broom, 1909) Morales & Shishkin, 2002 [Capitosaurus africanus Broom, 1909; Parotosaurus africanus (Broom, 1909) Romer, 1947; Parotosuchus africanus (Broom, 1909) Chernin, 1978; Wellesaurus africanus (Broom, 1909) Demiani, 2001]
†Wetlugasaurus Ryabinin, 1930
†W. angustifrons Ryabinin, 1930
†W. groenlandicus Säve-Söderbergh, 1935
†W. malachovi Novikov, 1990
†W. samarensis Sennikov, 1981
Reference(s):
- Milner, A. R., 1994: Late Triassic and Jurassic amphibians: Fossil record and phylogeny. 5-23
in Fraser, N. C. & Sues, H-D. 1994: In the Shadow of the Dinosaurs.
–Cambridge University Press, New York. 1994 - Morales, M. & Shishkin, M. A., 2002: A re-assessment of Parotosaurus africanus (Broom), a capitosauroid temnospondyl
amphibian from the Triassic of South Africa.
–Journal of Vertebrate Paleontology: Vol. 22, #1, pp. 1-11 - Schoch, R. R., 2000: The origin and interrelationships of Triassic capitosaurid amphibians.
–Paleontology: Vol. 43, #4, pp. 705-727 - Sulej, T. & Nied´zwiedzki, R., 2013: a new large capitosaurid temnospondyl amphibian from the Early Triassic of Poland.
–Acta Palaeontologica Polonica: Vol. 58, #1, pp. 65-75 [doi: 10.4202/app.2011.0025]