After Froese & Pauly, 2001 [“The FishBase”]
<==o Bothidae () [N: 20/115] [FB: 20/132]
|-- †Eobothus minimus M. Eoc. As. Eu.
|-- †Evesthes Mioc. NA.
|-- †Imhoffius M. Eoc. Eu.
|-- Arnoglossus Mioc. Eu. R. Oc. [25]
|--o Asterorhombus Tanaka, 1915 [7]
| |-- A. cocosensis (Bleeker, 1855) [taxonomic history] (Cocos Island flounder)
| |-- A. filifer Hensley & Randall, 2003
| `-- A. intermedius (Bleeker, 1865) fide Amaoka, 1969 [taxonomic history] (intermediate flounder)
|--o Bothus Eoc. Olig. Eu. R. Oc. [13]
| |-- B. assimilis (Günther, 1862)
| |-- B. constellatus (Jordan, 1889) (Pacific eyed flounder)
| |-- B. guibei Stauch, 1966 (Guinean flounder)
| |-- B. leopardinus (Günther, 1862) (Pacific leopard flounder)
| |-- B. lunatus (Linnaeus, 1758) (Plate fish)
| |-- B. maculiferus (Poey, 1860) (Mottled flounder)
| |-- B. mancus (Broussonet, 1782) (Flowery flounder)
| |-- B. mellissi Norman, 1931 (St. Helena flounder)
| |-- B. myriaster (Temminck & Schlegel, 1846) (Indo-Pacific oval flounder)
| |-- B. ocellatus (Agassiz, 1831) (Eyed flounder)
| |-- B. pantherinus (Rüppell, 1830) (Leopard flounder)
| |-- B. podas (Delaroche, 1809) (Wide-eyed flounder)
| `-- B. robinsi Topp & Hoff, 1972 (Twospot flounder)
|--o Chascanopsetta [Pelicanichthys] [5]
| |-- C. crumenalis (Gilbert & Cramer, 1897)
| |-- C. lugubris Alcock, 1894 (Pelican flounder)
| |-- C. micrognathus Amaoka & Yamamoto, 1984
| |-- C. prognathus Norman, 1939
| `-- C. prorigera Gilbert, 1905
|--o Crossorhombus [4]
| |-- C. azureus (Alcock, 1889) (Blue flounder)
| |-- C. howensis Hensley & Randall, 1993 (Lord Howe Island flounder)
| |-- C. kobensis (Jordan & Starks, 1906) (Kobe flounder)
| `-- C. valderostratus (Alcock, 1890) (Broadbrow flounder)
|--o Engyophrys [2]
| |-- Engyophrys sanctilaurentia Jordan & Bollman, 1890 (Speckled-tail flounder)
| `-- Engyophrys senta Ginsburg, 1933 (American spiny flounder)
|-- Engyprosopon [Scaeops] [25]
|--o Grammatobothus [3]
| |-- G. krempfi Chabanaud, 1929 (Krempf's flounder)
| |-- G. pennatus (Ogilby, 1913) (Pennant flounder)
| `-- G. polyophthalmus (Bleeker, 1865) )Threespot flounder)
|-- Japonolaeops dentatus Amaoka, 1969
|-- Kamoharaia megastoma (Kamohara, 1936) (Wide-mouthed flounder)
|--o Laeops [9]
| |-- L. clarus Fowler, 1934 (Clear fin-base flounder)
| |-- L. cypho Fowler, 1934
| |-- L. gracilis Fowler, 1934 (Philippine slender flounder)
| |-- L. guentheri Alcock, 1890 (Günther's flounder)
| |-- L. kitaharae (Smith & Pope, 1906)
| |-- L. natalensis Norman, 1931 (Khaki flounder)
| |-- L. nigromaculatus von Bonde, 1922 (Blackspotted flounder)
| |-- L. parviceps Günther, 1880 (Small headed flounder)
| `-- L. pectoralis (von Bonde, 1922) (Longarm flounder)
|-- Lophonectes gallus Günther, 1880 (Crested flounder)
|--o Monolene [9]
| |-- M. antillarum Norman, 1933 (Slim flounder)
| |-- M. asaedai Clark, 1936 (Asaedae flounder)
| |-- M. atrimana Goode & Bean, 1886
| |-- M. dubiosa Garman, 1899 (Acapulco flounder)
| |-- M. helenensis Amaoka & Imamura, 2000
| |-- M. maculipinna Garman, 1899 (Pacific deepwater flounder)
| |-- M. mertensi (Poll, 1959) (Merten's moonflounder)
| |-- M. microstoma Cadenat, 1937 (Smallmouth moonflounder)
| `-- M. sessilicauda Goode, 1880 (Deepwater flounder)
|-- Neolaeops microphthalmus (von Bonde, 1922) (Crosseyed flounder)
|--o Parabothus [9]
| |-- P. amaokai Parin, 1983
| |-- P. budkeri (Chabanaud, 1942)
| |-- P. chlorospilus (Gilbert, 1905)
| |-- P. coarctatus (Gilbert, 1905)
| |-- P. filipes Amaoka, Mihara & Rivaton, 1997
| |-- P. kiensis (Tanaka, 1918)
| |-- P. malhensis (Regan, 1908)
| |-- P. polylepis (Alcock, 1889) (Manyscaled flounder)
| `-- P. taiwanensis Amaoka & Shen, 1993
|-- Perissias taeniopterus (Gilbert, 1890) (Striped-fin flounder)
|--o Psettina [9]
| |-- P. brevirictis (Alcock, 1890)
| |-- P. filimana Li & Wang, 1982
| |-- P. gigantea Amaoka, 1963 (Rough-scaled flounder)
| |-- P. hainanensis (Wu & Tang, 1935)
| |-- P. iijimae (Jordan & Starks, 1904)
| |-- P. profunda (Weber, 1913)
| |-- P. senta Amaoka & Larson, 1999
| |-- P. tosana Amaoka, 1963
| `-- P. variegata (Fowler, 1934)
|--o Taeniopsetta [2]
| |-- T. ocellata (Günther, 1880) (Indo-Pacific ocellated flounder)
| `-- T. radula Gilbert, 1905
|--o Tosaramphus [5]
| |-- T. brevis Amaoka, Mihara & Rivaton, 1997
| |-- T. longimanus Amaoka, Mihara & Rivaton, 1997
| |-- T. neocaledonicus Amaoka & Rivaton, 1991
| |-- T. nielseni Amaoka & Rivaton, 1991
| `-- T. octoculatus Amaoka, 1969
`--o Trichopsetta [4]
|-- T. caribbaea Anderson & Gutherz, 1967 (Caribbean flounder)
|-- T. melasma Anderson & Gutherz, 1967
|-- T. orbisulcus Anderson & Gutherz, 1967
`-- T. ventralis (Goode & Bean, 1885) (Sash flounder)
Reference(s):
- Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698.
- Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698.
- Frickhinger, K. A., 1995: Fossil Atlas – Fishes.
–Mergus – Publishers for Natural History and Pet Books, Hans A. Baensch, Malle, Germany, 1-1088
- Froese, R. & Pauly, D. (eds), 2001: FishBase.
–iNet: www.fishbase.org
- Hensley, D. A., 2005: Revision of the genus Asterorhombus (Pleuronectiformes: Bothidae).
–Copeia: Vol. 2005 [105], #3, pp. 445-460
- Hensley, D. A., 2003: On the Status of Rhombus cocosensis Bleeker, and Description of a New Species of Engyprosopon Based on Misidentification of This Species.
–Copeia: Vol. 2003 [103], #4, pp. 833-837
- Lahti, S., Malmström, K. K., Koli, L., Leikola, A., Syrjämäki, J. & Lahti, J., 1980: Zoo, Suuri Eläinkirja 5: Kalat, sammakkoeläimet, matelijat.
–Werner Söderström Osakeyhtiö, Porvoo-Helsinki-Juva, 1980.
Original work [in French]: Beauté Du Monde Animal: IX Reptiles/Amphibiens & X Poissons.
–Rizzoli Editore, Milano, 1968,
- Nelson, J. S., 1994: Fishes of the world.
–John Wiley & Sons Inc., New York, 1994, xx-600