After Froese & Pauly, 2001 ["The FishBase"], Kinziger, Wood & Welsh, 2001, Page, Hartman & Near, 2003 and Powers & Mayden, 2003
<==o Etheostoma (darters; nuoliahvenet) [116 species] |--o (Catonotus) | |-- E. (C.) oophylax Ceas & Page, 1992 (Guardian darter) | `--+-- E. (C.) squamiceps Jordan, 1877 (Spottail darter) | `--+--+-- E. (C.) percnurum Jenkins, 1994 | | `-- E. (C.) flabellare Rafinesque, 1819 (Fantail darter) | `--+-- E. (C.) kennicotti (Putnam, 1863) (Stripetail darter) | `--+--+-- E. (C.) barbouri Kuehne & Small, 1971 (Teardrop darter) | | `-- E. (C.) basilare Page, Hartman & Near, 2003 (Corrugated darter) | `--+-- E. (C.) virgatum (Jordan, 1880) (Striped darter) | `--+-- E. (C.) obeyense Kirsch, 1892 (Barcheek darter) | `--+-- E. (C.) derivativum Page, Hartman & Near, 2003 (Stone darter) | `--+-- E. (C.) striatulum Page & Braasch, 1977 (Striated darter) | `-- E. (C.) smithi Page & Braasch, 1976 (Slabrock darter) |--o (Ulocentra) [18] | |-- E. (U.) coosae (Fowler, 1945) (Coosa darter) | |--+-- E. (U.) cervus Powers & Mayden, 2003 (Chickasaw darter) | | `-- E. (U.) pyrrhogaster Bailey & Etnier, 1988 (Firebelly darter) | |--o E. (U.) simoterum -group | | |-- E. (U.) baileyi Page & Burr, 1982 (Emerald darter) | | |-- E. (U.) barrenense Burr & Page, 1982 (Splendid darter) | | |-- E. (U.) rafinesquei Burr & Page, 1982 (Kentucky darter) | | `-- E. (U.) simoterum (Cope, 1868) (Snubnose darter) | `?-o E. (U.) duryi -group | |-- E. (U.) bellator | |-- E. (U.) brevirostrum Suttkus & Etnier, 1991 (Holiday darter) | |-- E. (U.) chermocki | |-- E. (U.) chlorosomum (Hay, 1881) (Bluntnose darter) | |-- E. (U.) duryi Henshall, 1889 (Black darter) | |-- E. (U.) etnieri Bouchard, 1977 (Cherry darter) | |-- E. (U.) flavum Etnier & Bailey, 1989 (Saffron darter) | |-- E. (U.) lachneri | |-- E. (U.) ramseyi | |-- E. (U.) raneyi | `-- E. (U.) scotti |--o E. variatum -group | |-- E. variatum Kirtland, 1838 (Variegate darter) | `--+-- E. tetrazonum (Hubbs & Black, 1940) (Missouri saddled darter) | `-- E. sp. cf. E. tetrazonum [Switzer & Wood, 2002] (Meramec River darter) |--o (Oligocephalus) | |-- E. spectabile (Agassiz, 1854) (Orangethroat darter) | `--+-- E. podostemone Jordan & Jenkins, 1889 (Riverweed darter) | `-- E. vitreum (Cope, 1870) (Glassy darter) `--+-- Percina `--+--+-- E. beanii | `-- E. calarum `--o (Nothonotus) |?- E. (N.) wapiti Etnier & Williams, 1989 (Boulder darter) |--+-- E. (N.) moorei Raney & Suttkus, 1964 (Yellowcheek darter) | `-- E. (N.) rubrum Raney & Suttkus, 1966 (Bayou darter) `--+-- E. (N.) rufilineatum (Cope, 1870) (Redline darter) `--+-- E. (N.) tippecanoe Jordan & Evermann, 1890 (Tippecanoe darter) `-- E. (N.) denoncourti Stauffer & van Snik, 1997Etheostoma -species I have not yet been able to assing in subgenera:
E. acuticeps Bailey, 1959 (Sharphead darter)
E. aquali Williams & Etnier, 1978 (Coppercheek darter)
E. asprigene (Forbes, 1878) (Mud darter)
E. australe Jordan, 1889 (Conchos darter)
E. bellum Zorach, 1968 (Orangefin darter)
E. bison Ceas & Page, 1997 (Buffalo darter)
E. blennioides Rafinesque, 1819 (Greenside darter)
E. blennius Gilbert & Swain, 1887 (Blenny darter)
E. boschungi Wall & Williams, 1974 (Slackwater darter)
E. burri Ceas & Page, 1997 (Brook Darter)
E. caeruleum Storer, 1845 (Rainbow darter)
E. camurum (Cope, 1870) (Bluebreast darter)
E. chienense Page & Ceas, 1992 (Relict darter)
E. chlorobranchium Zorach, 1972 (Greenfin darter)
E. cinereum Storer, 1845 (Ashy darter)
E. collettei Birdsong & Knapp, 1969 (Creole darter)
E. collis (Hubbs & Cannon, 1935) (Carolina darter)
E. corona Page & Ceas, 1992 (Crown darter)
E. cragini Gilbert, 1885 (Arkansas darter)
E. crossopterum Braasch & Mayden, 1985 (Fringed darter)
E. davisoni Hay, 1885 (Choctawhatchee darter)
E. ditrema Ramsey & Suttkus, 1965 (Coldwater darter)
E. edwini (Hubbs & Cannon, 1935) (Brown darter)
E. euzonum (Hubbs & Black, 1940) (Arkansas saddled darter)
E. exile (Girard, 1859) (Iowa darter)
E. fonticola (Jordan & Gilbert, 1886) (Fountain darter)
E. forbesi Page & Ceas, 1992 (Barrens darter)
E. fricksium Hildebrand, 1923 (Savannah darter)
E. fusiforme (Girard, 1854) (Swamp darter)
E. gracile (Girard, 1859) (Slough darter)
E. grahami (Girard, 1859) (Rio Grande darter)
E. histrio Jordan & Gilbert, 1887 (Harlequin darter)
E. hopkinsi (Fowler, 1945) (Christmas darter)
E. inscriptum (Jordan & Brayton, 1878) (Turquoise darter)
E. jessiae (Jordan & Brayton, 1878) (Blueside darter)
E. jordani Gilbert, 1891 (Greenbreast darter)
E. juliae Meek, 1891 (Yoke darter)
E. kanawhae (Raney, 1941) (Kanawha darter)
E. kantuckeense Ceas & Page, 1997 (Highland Rim Darter)
E. lepidum (Baird & Girard, 1853) (Greenthroat darter)
E. longimanum Jordan, 1888 (Longfin darter)
E. lugoi Norris & Minckley, 1997 (Tufa darter)
E. luteovinctum Gilbert & Swain, 1887 (Redband darter)
E. lynceum Hay, 1885 (Brighteye darter)
E. maculatum Kirtland, 1841 (Spotted darter)
E. mariae (Fowler, 1947) (Pinewoods darter)
E. microlepidum Raney & Zorach, 1967 (Smallscale darter)
E. microperca Jordan & Gilbert, 1888 (Least darter)
E. neopterum Howell & Dingerkus, 1978 (Lollipop darter)
E. nianguae Gilbert & Meek, 1887 (Niangua darter)
E. nigripinne Braasch & Mayden, 1985 (Blackfin darter)
E. nigrum Rafinesque, 1820 (Johnny darter)
E. nuchale Howell & Caldwell, 1965 (Watercress darter)
E. okaloosae (Fowler, 1941) (Okaloosa darter)
E. olivaceum Braasch & Page, 1979 (Sooty darter)
E. olmstedi Storer, 1842 (Tessellated darter)
E. osburni (Hubbs & Trautman, 1932) (Candy darter)
E. pallididorsum Distler & Metcalf, 1962 (Paleback darter)
E. parvipinne Gilbert & Swain, 1887 (Goldstripe darter)
E. perlongum (Hubbs & Raney, 1946) (Waccamaw darter)
E. pottsii (Girard, 1859) (Chihuahua darter)
E. proeliare (Hay, 1881) (Cypress darter)
E. pseudovulatum Page & Ceas, 1992 (Egg-mimic darter)
E. punctulatum (Agassiz, 1854) (Stippled darter)
E. radiosum (Hubbs & Black, 1941) (Orangebelly darter)
E. rupestre Gilbert & Swain, 1887 (Rock darter)
E. sagitta (Jordan & Swain, 1883) (Arrow darter)
E. saludae (Hubbs & Cannon, 1935) (Saluda darter)
E. sanguifluum (Cope, 1870) (Bloodfin darter)
E. segrex Norris & Minckley, 1997 (Río Salado darter)
E. sellare (Radcliffe & Welsh, 1913) (Maryland darter)
E. serrifer (Hubbs & Cannon, 1935) (Sawcheek darter)
E. stigmaeum (Jordan, 1877) (Speckled darter)
E. swaini (Jordan, 1884) (Gulf darter)
E. swannanoa Jordan & Evermann, 1889 (Swannanoa darter)
E. tallapoosae Suttkus & Etnier, 1991
E. tecumsehi Ceas & Page , 1997 (Shawnee darter)
E. thalassinum (Jordan & Brayton, 1878) (Seagreen darter)
E. trisella Bailey & Richards, 1963 (Trispot darter)
E. tuscumbia Gilbert & Swain, 1887 (Tuscumbia darter)
E. vulneratum (Cope, 1870) (Wounded darter)
E. whipplii (Girard, 1859) (Redfin darter)
E. zonale (Cope, 1868) (Banded darter)
E. zonifer (Hubbs & Cannon, 1935) (Backwater darter)
E. zonistium Bailey & Etnier, 1988 (Bandfin darter)
Reference(s):
- Froese, R. & Pauly, D. (eds), 2001: FishBase.
–iNet: www.fishbase.org - Kinziger, A. P., Wood, R. M. & Welsh, S. A., 2001: Systematics of Etheostoma tippecanoe and Etheostoma denoncourti (Perciformes: Percidae).
–Copeia: Vol. 101, #1, pp. 235-239 - Page, L. M., Hartman, M. & Near, T., 2003: Phylogenetic Relationships of Barcheek Darters (Percidae: Etheostoma, Subgenus Catonotus) with Descriptions of two new species.
–Copeia: Vol. 103, #3, pp. 512-530 - Powers, S. L. & Mayden, R. L., 2003: Etheostoma cervus: A New Species from the Forked Deer River System in Western Tennessee with Comparison to Etheostoma pyrrhogaster (Percidae: subgenus Ulocentra).
–Copeia: Vol. 103, #3, pp. 576-582 - Switzer, J. F. & Wood, R. M., 2002: Molecular systematics and historical biogeography of the Missouri Saddled Darter Etheostoma tetrazonum (Actinopterygii: Percidae).
–Copeia: Vol. 102, #2, pp. 450-455