After Carroll, 1988, Nelson, 1994, and Riva Rossi, Gosztonyi & Cozzuol, 2000, with Recent species from Froese & Pauly, 2001 [“The Fishbase”]

Note:

While naming a new fossil cusk-eel species, †Genypterus valdensensis, Riva Rossi et al. mention that it belongs to a tribe Lepophidiini. This and other tribes were not mentioned in Nelson (1994), which is the main source of this and other fish-pages. Also the FishBase does not use subfamilies or tribes, which makes the combination of it and Nelson (1994) much more difficult.

<==o Ophidiinae Rafinesque, 1810 (partanilkat) [8/60]
   |-- †Preophidion Eoc. NA.
   |?- Grimaldichthys profundissimus (grimaldikala)
   |--o ?Lepophidiini Robins, 1961
   |  |-- Cherublemma emmelas (Gilbert, 1890) (Black brotula)
   |  |--o Genypterus Philippi, 1857 (cusk-eels) [8]
   |  |  |-- †G. valdensensis Riva Rossi et al., 2000
   |  |  |-- G. tigerinus Klunzinger, 1872 (Rock ling)
   |  |  |-- G. nigricans Philippi, 1857
   |  |  |--+-- G. blacodes (Forster, 1801) (Pink cusk-eel)
   |  |  |  |?- G. microstomus Regan, 1901
   |  |  |  `?- G. capensis (Smith, 1847) (Kingklip)
   |  |  |-- G. brasiliensis Regan, 1903
   |  |  |-- G. chilensis (Guichenot, 1848) (Red cusk-eel)
   |  |  `-- G. maculatus (Tschudi, 1846) (Black cusk-eel)
   |  `--o Lepophidium [13]
   |     |-- L. aporrhox Robins, 1961 (Dusky cusk-eel)
   |     |-- L. brevibarbe (Cuvier, 1829) (Shortbeard cusk-eel)
   |     |-- L. jeannae Fowler, 1941 (Mottled cusk-eel)
   |     |-- L. kallion Robins, 1959
   |     |-- L. marmoratum (Goode & Bean, 1885)
   |     |-- L. microlepis (Gilbert, 1890) (Silver cusk eel)
   |     |-- L. negropinna Hildebrand & Barton, 1949 (Specklefin cusk eel)
   |     |-- L. pardale (Gilbert, 1890) (Leopard cusk eel)
   |     |-- L. pheromystax Robins, 1960 (Blackedge cusk-eel)
   |     |-- L. profundorum (Gill, 1863) (Blackrim cusk-eel)
   |     |-- L. prorates (Jordan & Bollman, 1890) (Prowspine cusk eel)
   |     |-- L. staurophor Robins, 1958 (Barred cusk-eel)
   |     `-- L. stigmatistium (Gilbert, 1890) (Blotchfin cusk eel)
   `--o “Ophidiini”
      |-- Chilara taylori (Girard, 1858) (Spotted cusk-eel)
      |-- Ophidion [Ophidium, Rissola] ?Paleoc. M. Eoc. Eu. Eoc. WInd. Mioc. Aus. R. Oc. (partanilkat) [23]
      |--o Otophidium [4]
      |  |-- O. chickcharney Böhlke & Robins, 1959 (Ghost cusk-eel)
      |  |-- O. dormitator Böhlke & Robins, 1959 (Sleeper cusk-eel)
      |  |-- O. indefatigabile Jordan & Bollman, 1890 (Bighead cusk eel)
      |  `-- O. omostigma (Jordan & Gilbert, 1882) (Polka-dot cusk-eel)
      |--o Parophidion [2]
      |  |-- P. schmidti (Woods & Kanazawa, 1951) (Dusky cusk-eel)
      |  `-- P. vassali (Risso, 1810)
      `-- Raneya brasiliensis (Kaup, 1856)

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