After Nelson, 1994, and Fang, 2003, with recent species from Froese & Pauly, 2001 [“The FishBase”]
<==o Rasborinae [Danioninae] (rasboras; rasborat)
|--o Amblypharyngodon U. Tert. NAs. R.
| |-- A. atkinsonii (Blyth, 1860) (Burmese carplet)
| |-- A. chakaiensis Babu Rao & Nair, 1978 (Veli Lake carplet)
| |-- A. chulabhornae Vidthayanon & Kottelat, 1990
| |-- A. melettinus (Valenciennes, 1844) (Attentive carplet)
| |-- A. microlepis (Bleeker, 1853) (Indian carplet)
| `-- A. mola (Hamilton, 1822) (Mola carplet)
|--o Aspidoparia
| |-- A. jaya (Hamilton, 1822) (Jaya)
| `-- A. morar (Hamilton, 1822)
|-- Barilius Hamilton, 1822
|--o Boraras
| |-- B. brigittae (Vogt, 1978) (hehkurasbora)
| |-- B. maculatus (Duncker, 1904) [Rasbora maculata] (kääpiöraspora; Dwarf rasbora)
| |-- B. merah (Kottelat, 1991) (minirasbora)
| |-- B. micros Kottelat & Vidthayanon, 1993
| `-- B. urophthalmoides (Kottelat, 1991) (juovaminirasbora, huutomerkkirasbora; Exclamation-point rasbora)
|-- Engraulicypris sardella (Günther, 1868) (Lake sardine)
|-- Macrochirichthys macrochirus (Valenciennes, 1844)
|--o Mesobola
| |-- M. brevianalis (Boulenger, 1908) (River sardine)
| `-- M. spinifer (Bailey & Matthes, 1971) (Malagarasi sardine)
|--o Neobola
| |-- N. bottegoi Vinciguerra, 1895
| |-- N. bredoi (Poll, 1945)
| |-- N. fluviatilis (Whitehead, 1962) (Athi sardine)
| |-- N. moeruensis (Boulenger, 1915)
| `-- N. stellae (Worthington, 1932)
|--o Oxygaster
| |-- O. anomalura Van Hasselt, 1823
| `-- O. pointoni (Fowler, 1934)
|?-o Pseudorasbora Mioc. R. As.
| |-- †P. macrocephala
| |-- P. elongata Wu, 1939
| |-- P. parva (Temminck & Schlegel, 1846) (stone moroko)
| `-- P. pumila Miyadi, 1930 (moroko, moroco)
|--+?-o Opsariichthys
| | |-- O. bidens Günther, 1873
| | `--o O. uncirostris (Temminck & Schlegel, 1846)
| | |-- O. u. uncirostris (Temminck & Schlegel, 1846)
| | `-- O. u. amurensis Berg, 1932 (Triplelip)
| |?- Opsarius koratensis (Smith, 1931)
| |--o Leptocypris Boulenger, 1900
| | |-- L. crossensis Howes & Teugels, 1989
| | |-- L. guineensis (Daget, 1962)
| | |-- L. konkoureensis Howes & Teugels, 1989
| | |-- L. lujae (Boulenger, 1909)
| | |-- L. modestus Boulenger, 1900
| | |-- L. niloticus (Joannis, 1835)
| | |-- L. taiaensis Howes & Teugels, 1989
| | |-- L. weeksii (Boulenger, 1899) [Barilius weeksii Boulenger, 1899; Raiamas weeksii (Boulenger) Howes, 1980]
| | `-- L. weynsii (Boulenger, 1899)
| |--o Opsaridium Peter, 1854
| | |-- O. engrauloides (Nichols, 1923)
| | |-- O. leleupi (Matthes, 1965)
| | |-- O. loveridgii (Norman, 1922)
| | |-- O. maculicauda (Pellegrin, 1926)
| | |-- O. microcephalum (Günther, 1864)
| | |-- O. microlepis (Günther, 1864) (Lake salmon)
| | |-- O. splendens Taverne & De Vos, 1997
| | |-- O. tweddleorum Skelton, 1996
| | |-- O. ubangiensis (Pellegrin, 1901)
| | `-- O. zambezense (Peters, 1852) (Barred minnow)
| `--o Raiamas Jordan, 1919
| |-- R. ansorgii (Boulenger, 1910)
| |-- R. batesii (Boulenger, 1914)
| |-- R. bola (Hamilton, 1822) (Trout barb)
| |-- R. buchholzi (Peters, 1876)
| |-- R. christyi (Boulenger, 1920) [Barilius christyi Boulenger, 1920; Opsaridium christyi (Boulenger) Howes, 1980] (Coppernose barb; kultasuuopsari)
| |-- R. guttatus (Day, 1870) (Burmese trout)
| |-- R. kheeli Stiassny, Schelly & Schliewen, 2006
| |-- R. levequei Howes & Teugels, 1989
| |-- R. longirostris (Boulenger, 1902)
| |-- R. macrostoma (Boulenger, 1913)
| |-- R. moorii (Boulenger, 1900) (Lake Rukwa minnow)
| |-- R. nigeriensis (Daget, 1959)
| |-- R. salmolucius (Nichols & Griscom, 1917)
| |-- R. scarciensis Howes & Teugels, 1989
| |-- R. senegalensis (Steindachner, 1870) (Silver fish)
| |-- R. shariensis (Fowler, 1949)
| `-- R. steindachneri (Pellegrin, 1908)
|--+-- Rasbora Tert. R. EInd
| `-- Rasboroides vaterifloris (Deraniyagala, 1930) [Rasbora vaterifloris Deraniyagala, 1930; Rasbora nigromarginata Meinken, 1957] (Pearly rasbora; helmiäisrasbora)
|-- Rastrineobola argentea (Pellegrin, 1904) (Silver cyprinid)
|--o Salmostoma
| |-- S. acinaces (Valenciennes, 1844) (Silver razorbelly minnow)
| |-- S. bacaila (Hamilton, 1822) (Large razorbelly minnow)
| |-- S. balookee (Sykes, 1839) (Bloch razorbelly minnow)
| |-- S. boopis (Day, 1874) (Boopis razorbelly minnow)
| |-- S. horai (Silas, 1951) (Hora razorbelly minnow)
| |-- S. kardahiensis Reddiah, 1980
| |-- S. novacula (Valenciennes, 1844) (Novacula razorbelly minnow)
| |-- S. orissaensis Banarescu, 1968 (Orissa razorbelly minnow)
| |-- S. panjabensis (Day, 1872) (Punjab razorbelly minnow)
| |-- S. phulo (Hamilton, 1822) (Finescale razorbelly minnow)
| |-- S. sardinella (Valenciennes, 1844) (Sardinella razorbelly minnow)
| |-- S. sladoni (Day, 1870)
| `-- S. untrahi (Day, 1869) (Mahanadi razorbelly minnow)
|--o Tanichthys Lin, 1932 (kardinaalikalat)
| |-- T. albonubes Lin, 1932 (kardinaalikala; White cloud mountain minnow)
| `-- T. micagemmae Freyhof & Herder, 2001 (juvakardinaalikala)
|--o Trigonostigma Kottelat & Witte, 1999 (kiilakyljet)
| |-- T. hengeli (Meinken, 1956) [Rasbora hengeli Meinken, 1956] (Glowlight rasbora; loistokiilakylki, pikkukiilakylki)
| |-- T. heteromorpha (Duncker, 1904) (Harlequin rasbora; kiilakylki)
| |-- T. espei (Meinken, 1967) [Rasbora heteromorpha espei Meinken, 1967] (Lambchop rasbora; rusokiilakylki)
| `-- T. somphongsi (Meinken, 1958)
|--o Zacco
| |-- Z. chengtui Kimura, 1934
| |-- Z. pachycephalus (Günther, 1868)
| |-- Z. platypus (Temminck & Schlegel, 1846) (Freshwater minnow)
| |-- Z. taiwanensis Chen, 1982
| `-- Z. temmincki (Temminck & Schlegel, 1846) (Dark chub)
`--o Danionini sensu Fang, 2003 (danionines; seeprakalat)
|--+--o Microdevario [Microrasbora in partim]
| | |-- M. gatesi (Herre, 1939) [Microrasbora gatesi Herre, 1939]
| | `--+-- M. kubotai (Kottelat & Witte, 1999) [Microrasbora kubotai Kottelat & Witte, 1999] (smaragdirasbora)
| | `-- M. nanus (Kottelat & Witte, 1999) [Microrasbora nana Kottelat & Witte, 1999]
| |--+--o Inlecypris Howes, 1980 [=? Devario]
| | | |-- I. jayarami (Barman, 1984) [Brachydanio jayarami (Barman, 1984)]
| | | `-- I. auropurpureus (Annandale, 1918) [Barilius auropurpureus Annandale, 1918; Devario auropurpureus (Annandale, 1918)]
| | |--o Laubuca Bleeker, 1860
| | | |-- L. caeruleostigmata H. M. Smith, 1931 [Chela caeruleostigmata (Smith, 1931)] (Leaping barb; sinikölibarbi)
| | | |-- L. dadiburjori Menon, 1952 [Chela dadidurjori (Menon, 1952)] (Dadio; hehkukölibarbi)
| | | |-- L. fasciata (Silas, 1958) [Chela fasciata Silas, 1958] (kiiltokölibarbi)
| | | |-- L. insularis Pethiyagoda, Kottelat, Anjana Silva, Maduwage & Meegaskumbura, 2008
| | | |-- L. lankensis (Deraniyagala, 1960)
| | | |-- L. laubuca (Hamilton, 1822) [Chela laubuca (Hamilton, 1822)] (Indian glass barb)
| | | |-- L. ruhuna Pethiyagoda, Kottelat, Anjana Silva, Maduwage & Meegaskumbura, 2008
| | | |-- L. varuna Pethiyagoda, Kottelat, Anjana Silva, Maduwage & Meegaskumbura, 2008
| | | `-- L. maassi (Weber & de Beaufort, 1912) [Chela maassi (Weber & de Beaufort, 1912)]
| | `--o Chela Hamilton, 1822; Eoc. Eu. R. As. EInd. (kölibarbit)
| | |-- C. cachius (F. Hamilton, 1822) (kölibarbi)
| | `-- C. khujairokensis Arunkumar, 2000
| `==o Devario Heckel, 1843 [Danio Hamilton, 1822 in partim; Paradanio Day, 1865; Eustira Günther, 1868; Rambaibarnia Fowler, 1934; Danioides Chu, 1935; Parabarilius Pellegrin & P. W. Fang, 1940; Daniops Smith, 1945; Brachydanio Weber & de Beaufort, 1916]
| |?- D. acrostomus (F. Fang & Kottelat, 1999) [Danio acrostomus Fang & Kottelat, 1999]
| |?- D. acuticephala (Hora, 1921)
| |?- D. aequipinnatus (McClelland, 1839) {Danio aequipinnatus (McClelland, 1839); D. malabaricus] (siniseeprakala; Giant danio)
| |?- D. affinis (Blyth, 1860)
| |?- D. annandalei (Chaudhuri, 1908) [Danio annandalei Chaudhuri, 1908]
| |?- D. anomalus Conway, Mayden & K. L. Tang, 2009
| |?- D. apopyris (F. Fang & Kottelat, 1999) [Danio apopyris Fang & Kottelat, 1999]
| |?- D. assamensis (Barman, 1984) [Danio assamensis Barman, 1984]
| |?- D. auropurpureus (Annandale, 1918)
| |?- D. browni (Regan, 1907)
| |?- D. fangfangae (Kottelat, 2000) [Danio fangfangae Kottelat, 2000]
| |?- D. fraseri (Hora, 1935) [Danio fraseri Hora, 1935]
| |?- D. gibber (Kottelat, 2000) [Danio gibber Kottelat, 2000]
| |?- D. horai (Barman, 1983)
| |?- D. laoensis (Pellegrin & P. W. Fang, 1940) [Parabarilius laoensis Pellegrin & Fang, 1940; Danio laoensis (Pellegrin & Fang, 1940)]
| |?- D. leptos (F. Fang & Kottelat, 1999) [Danio leptos Fang & Kottelat, 1999]
| |?- D. manipurensis (Barman, 1987)
| |?- D. naganensis (Chaudhuri, 1912) [Danio naganensis Chaudhuri, 1912]
| |?- D. neilgherriensis (F. Day, 1867) [Danio neilgherriensis (Day, 1867)]
| |?- D. pathirana (Kottelat & Pethiyagoda, 1990) [Danio pathirana Kottelat & Pethiyagoda, 1990] (Barred danio; safiiriseeprakala)
| |?- D. peninsulae (H. M. Smith, 1945)
| |?- D. regina (Fowler, 1934) [Danio regina Fowler, 1934]
| |?- D. salmonata (Kottelat, 2000) [Danio salmonata Kottelat, 2000]
| |?- D. shanensis (Hora, 1928) (Hora danio)
| |?- D. sondhii (Hora & Mukerji, 1934) [Brachydanio sondhii (Hora & Mukerji, 1934)]
| |?- D. spinosus (F. Day, 1870) [Danio spinosus Day, 1870]
| |?- D. strigillifer (Myers, 1924)
| |?- D. suvatti (Fowler, 1939)
| |?- D. xyrops F. Fang & S. O. Kullander, 2009
| |?- D. yuensis (Arunkumar & Tombi Singh, 1998)
| |?- D. sp. “Giraffe”
| |-- D. devario (Hamilton, 1822) (Sind danio)
| `--+-- Betadevario ramachandrani Pramod, Fang, Rema Devi, Liao,, Indra, Jameela Beevi & Kullander, 2010
| `--+--+-- D. kakhienensis Anderson, 1879
| | `-- D. malabaricus (Jerdon, 1849) (Malabar danio; siniseeprakala)
| `--+-- D. chrysotaeniatus (Chu, 1981) [Danio chrysotaeniatus Chu, 1981]
| `--+-- D. maetaengensis (Fang, 1997) [Danio maetaengensis Fang, 1997]
| `--+-- D. interruptus (Day, 1870)
| `-- D. apogon (Chu, 1981) [Danio apogon Chu, 1981]
`--+--+--o Danio Hamilton, 1822 sensu stricto (zebrafishes, danios; seeprakalat)
| | |?- D. choprae Hora, 1928
| | |?- D. feegradei Hora, 1937 (sinikylkiseeprakala)
| | |?- D. jaintianensis (Sen, 2007)
| | |?- D. meghalayensis Sen & Dey, 1985
| | |?- D. muongthanhensis V. H. Nguyen, 2001
| | |?- D. nigrofasciatus (Day, 1870) (täpläseeprakala; Dwarf danio)
| | |?- D. quagga S. O. Kullander, Liao & Fang, 2009
| | |?- D. quangbinhensis (Nguyen, V. T. Le & X. K. Nguyen, 1999)
| | |?- D. shanensis Hora, 1928 (Hora danio)
| | |?- D. tinwini Kullander & Fang, 2009
| | |?- D. trangi Ngô, 2003
| | |-- D. dangila (Hamilton, 1822) Bleeker, 1862 [type]
| | `--+--+-- D. choprae Hora, 1928 [D. choprai] (rubiiniseeprakala)
| | | `--+-- D. erythromicron (Annandale) Kottelat & Witte, 1999; “Microrasbora” erythromicron Annandale, 1918] (raitarasbora)
| | | `-- D. margaritatus (Roberts, 2007) [Microraspora sp. “Galaxy” Celestichthys margaritatus Roberts, 2007] (celestial pearl danio; helmirasbora)
| | `--+--+-- D. aesculapii Kullander & F. Fang, 2009
| | | `--+-- D. kyathit Fang, 1998 (rusoseeprakala)
| | | `-- D. sp. “Burma” (Ozelot Danio; oselottiseeprakala)
| | `--+--o D. rerio (Hamilton, 1822) (seeprakala; Zebra danio)
| | | `-- D. rerio var. “frankei” [D. frankei (Meinken, 1963)] (Leopard danio; pilkkuseeprakala)
| | `--+-- D. kerri Smith, 1931 [Davario kerri (Smith, 1931); Brachydanio kerri (Smith, 1931)] (kultaseeprakala; hehkuseeprakala; Blue danio)
| | `--+-- D. roseus Fang & Kottelat, 2000
| | `--o D. albolineatus (Blyth, 1860) [Nuria albolineata Blyth, 1860; Brachydanio albolineatus (Blyth, 1860)] (hohtoseeprakala; Pearl danio)
| | |-- D. albolineatus var.. “pulcher”
| | `-- D. albolineatus var. “tweediei”
| `--o Esomus (lentobarbit)
| |-- E. ahli Hora & Mukerji, 1928
| |-- E. altus (Blyth, 1860)
| |-- E. barbatus (Jerdon, 1849) (South Indian flying barb)
| |-- E. caudiocellatus Ahl, 1923
| |-- E. danricus (Hamilton, 1822) (Flying barb)
| |-- E. lineatus Ahl, 1923 (Striped flying barb; juovalentobarbi)
| |-- E. longimanus (Lunel, 1881) (Mekong flying barb)
| |-- E. malayensis Ahl, 1923 (Malayan flying barb)
| |-- E. metallicus Ahl, 1923
| `-- E. thermoicos (Valenciennes, 1842)
`--+--+?-o Paedocypris Kottelat, Britz, Tan & Witte, 2005
| | |-- P. progenetica Kottelat, Britz, Tan & Witte, 2006
| | |-- P. micromegethes Kottelat, Britz, Tan & Witte, 2006
| | `-- P. carbunculus Britz & Kottelat, 2008
| |-- Fangfangia spinocleithralis Britz, Kottelat & Tan, 2012
| `--o Sundadanio Kottelat & Witte, 1999
| |-- S. axelrodi (Brittan, 1976) Kottelat & Witte, 1999 [Rasbora axelrodi Brittan, 1976] (safiirirasbora)
| |-- S. atomus Conway, Kottelat & Hui, 2011
| |-- S. gargula Conway, Kottelat & Hui, 2011
| |-- S. goblinus Conway, Kottelat & Hui, 2011
| |-- S. rubellus Conway, Kottelat & Hui, 2011
| |-- S. echinus Conway, Kottelat & Hui, 2011
| |-- S. retiarius Conway, Kottelat & Hui, 2011
| `-- S. margarition Conway, Kottelat & Hui, 2011
`--+--o Danionella Roberts, 1986
| |-- D. translucida Roberts, 1986 (lasirasbora)
| `-- D. mirifica Britz, 2003
`--o Microrasbora Annandale, 1918
|-- M. microphthalma Jiang, Chen & Yang, 2008
`-- M. rubescens Annandale, 1918
Reference(s):
- Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698.
- Carroll, R. L., 1988: Appendix. 594-648.
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698.
- Conway, K. W., Kottelat, M. &Sundadanio (Ostariophysi: Cyprinidae) with descriptions of seven new species from Indonesia and Malaysia.
–Ichthyological Exploration of Freshwaters: Vol. 22, #3, pp. 251-288
- Fang, F., 2003: Phylogenetic Analysis of the Asian Cyprinid Genus Danio (Teleostei, Cyprinidae).
–Copeia: Vol. 103, #4, pp. 714-728
- Frickhinger, K. A., 1995: Fossil Atlas – Fishes.
–Mergus – Publishers for Natural History and Pet Books, Hans A. Baensch, Malle, Germany, 1-1088
- Froese, R. & Pauly, D. (eds), 2001: FishBase.
–iNet: www.fishbase.org
- Lahti, S., Malmström, K. K., Koli, L., Leikola, A., Syrjämäki, J. & Lahti, J., 1980: Zoo, Suuri Eläinkirja 5: Kalat, sammakkoeläimet, matelijat.
–Werner Söderström Osakeyhtiö, Porvoo-Helsinki-Juva, 1980.
Original publication [in French]: Beauté Du Monde Animal: IX Reptiles/Amphibiens & X Poissons.
–Rizzoli Editore, Milano 1968.
- Long, J. A., 1995: The Rise of Fishes: 500 million years of evolution.
–Johns Hopkins University Press, Baltimore & London, pp. 1-223
- Nelson, J. S., 1994: Fishes of the world.
–John Wiley & Sons Inc., New York, 1994, xx-600
- Pramod, P. K., Fang, F., Rema Devi, K., Liao, T.-Y., Indra, T. J., Jameela Beevi, K. S. & Kullander, S. O., 2010: Betadevario ramachandrani, a new danionine genus and species from the Western Ghats of India (Teleostei: Cyprinidae: Danioninae).
–Zootaxa: Vol. #2519, pp. 31-47
- Roberts, T. R., 2007: The “celestial pearl danio”, a new genus ans species of colourful minute cyprinid fish from Myanmar (Pisces: Cypriniformes).
–The Raffles Bulletin of Zoology: Vol. 55, #1, pp. 131-140
- Stiassny, M. L. J., Schelly, R. C. & Schliewen, U. K., 2006: A new species of Raiamas (Teleostei: Cyprinidae) from the lower Congo river, with a phylogenetic assessment of the genetic limits of the predatory cyprinid genera Opsaridium, Raiamas, and Leptocypris.
–Copeia: Vol. 2006, #3, pp. 370-377
- Varjo, M., 2001: Akvaario-opas. [Aquarium Guide]
–AquaGeo & Gummerus Kirjapaino Oy, Jyväskylä, 2001, 1-143