After Broughton & Gold, 2000, with species from Froese & Pauly, 2001 [“The Fishbase”]
<==o Cyprinella Girard, 1856 ()? [30] |?- C. monacha (Cope, 1868) (Spotfin chub) [falls out of the monophyletic Cyprinella sensu Broughton & Gold, 2000] `--+-- C. callistia (Jordan, 1877) (Alabama shiner) `--+--+-- C. gibbsi (Howell & Williams, 1971) (Tallapoosa shiner) | `-- C. trichroistia (Jordan & Gilbert, 1878) (Tricolor shiner) `--+--+-- C. caerulea (Jordan, 1877) (Blue shiner) | `--+--+-- C. callisema (Jordan, 1877) (Ocmulgee shiner) | | `-- C. nivea (Cope, 1870) (Whitefin shiner) | `--+-- C. leedsi (Fowler, 1942) (Bannerfin shiner) | `--+-- C. callitaenia (Bailey & Gibbs, 1956) (Bluestripe shiner) | `-- C. zanema (Jordan & Brayton, 1878) (Santee chub) `--+--+-- C. xaenura (Jordan, 1877) (Altamaha shiner) | `--+-- C. labrosa (Cope, 1870) (Thicklip chub) | `-- C. pyrrhomelas (Cope, 1870) (Fieryblack shiner) `--+--+--+-- C. analostana Girard, 1859 (Satinfin shiner) | | `-- C. chloristia (Jordan & Brayton, 1878) (Greenfin shiner) | `--+-- C. proserpina (Girard, 1856) (Proserpine shiner) | `--+-- C. camura (Jordan & Meek, 1884) (Bluntface shiner) | `-- C. galactura (Cope, 1868) (Whitetail shiner) `--+--+-- C. venusta Girard, 1856 (Blacktail shiner) | `--+-- C. spiloptera (Cope, 1867) (Spotfin shiner) | `-- C. whipplii Girard, 1856 (Steelcolor shiner) `--+-- C. lepida Girard, 1856 (Plateau shiner) |-- C. lutrensis (Baird & Girard, 1853) [Leuciscus bubalinus Baird & Girard] (Red shiner) `-- C. formosa (Girard, 1856) (Beautiful shiner)
Cyprinella not treated by Broughton & Gold, 2000:
C. alvarezdelvillari Contreras-Balderas & Lourdes Lozano, 1994
C. bocagrande (Chernoff & Miller, 1982)
C. garmani (Jordan, 1885)
C. panarcys Hubbs & Miller, 1978 (Conchos shiner)
C. rutila (Girard, 1856)
C. xanthicara (Minckley & Lytle, 1969)
Reference(s):
- Broughton, R. E. & Gold, J. R., 2000: Phylogenetic relationships in the North American cyprinid genus Cyprinella (Actinopterygii: Cyprinidae) based on sequences of the mitochondrial ND2 and ND4L genes.
–Copeia: Vol. 100, #1, pp. 1-10 - Froese, R. & Pauly, D. (eds), 2001: FishBase.
–iNet: www.fishbase.org - Nelson, J. S., 1994: Fishes of the world.
–John Wiley & Sons Inc., New York, 1994, xx-600