After Šlechtová, Bohlen, Freyhof & Ráb, 2006 (fig. 5), with Recent species from Froese & Pauly, 2001 [“The Fishbase”] and Kottelat, 2012
<==o Botiidae Berg, 1940 (botiine loaches; loistonuoliaiset, botiat, “kirjonuoliaiset”) |--o Leptobotiinae | |--o Parabotia Dabry de Thiersant, 1872 | | |?- P. bimaculatus Chen, 1980 emend. Kottelat, 2012 [pro Parabotia bimaculata Chen, 1980] | | |?- P. brevirostris Zhu & Zhu, 2012 | | |?- P. curtus (Temminck & Schlegel, 1846) [Cobitis curta Temminck & Schlegel, 1846; Leptobotia curta (Temminck & Schlegel, 1846)] (Kissing loach) | | |?- P. dubius Kottelat, 2001 emend. Kottelat, 2012 [Botia elongata Mai, 1978; Parabotia dubia Kottelat, 2001; ?Parabotia kimluani Nguyen, 2005; ?Parabotia vancuongi Nguyen, 2005] | | |?- P. heterocheilus Zhu & Chen, in Ni & Wu, 2006 emend. Kottelat, 2012 [pro Parabotia heterocheilus Zhu & Chen, in Ni & Wu, 2006] | | |?- P. maculosus (Wu, 1939) [Botia maculosa] Wu, 1939 | | |?- P. mantschuricus Berg, 1907 [Leptobotia mantschurica Berg, 1907] (Manchurian spiny loach) | | |?- P. parvus Chen, 1980 | | |-- P. lijiangensis Chen, 1980 | | `--+-- P. kiangensis Liu & Guo, 1986 | | `--+-- P. banarescui (Nalbant, 1965) [Leptobotia banarescui Nalbant, 1965] | | `-- P. fasciatus Dabry de Thiersant, 1872 [Nemacheilus xanthi Günther, 1888; Botia multifasciata Regan, 1905; leptobotia intermedia Mori, 1929; Leptobotia hopeiensis Shaw & Tchang, 1931; Leptobotia kudoii Mori, 1933; Botia kwangensis Fang, 1936; Botia wui Chang, 1944] | `--o Leptobotia Bleeker, 1870 | |?- L.? cheni (Herre & Lin, 1936) [Nemacheilus cheni Herre & Lin, 1936] | |?- L.? variegata (Günther, 1889) [Botia variegata Günther, 1889] | |?- L. flavolineata Wang, 1981 | |?- L. guilinensis Chen, 1980 | |?- L. hengyangensis Huang & Zhang, 1986 | |?- L. microphthalma Fu & Ye, 1983 (Chinese Loach) | |?- L. orientalis Xu, Fang & Wang , 1984 | |?- L. posterodorsalis Lan & Chen, in Chen & Lan, 1992 | |?- L. punctata Li, Li & Chen, 2008 | |?- L. tientaienensis (Wu, 1930) [Botia tientaienensis Wu, 1930; Botia compressicauda Nichols, 1931; Botia tientaienensis hansuiensis Fang & Hsu, 1980] | |?= and numerous undescribed species? [The Loaches Online] | |-- L. guilinensis (puikkonuoliainen) | `--+-- L. elongata (Bleeker, 1870) [Botia elongata Bleeker, 1870; Cobitis variegata Dabry de Thiersant, 1872; Botia citrauratea Nichols, 1925] (Royal Clown Loach) | `--+-- L. pellegrini Fang, 1936 (Chinese Loach) | `--+-- L. rubrilabris Dabry de Thiersant, 1872 [Botia pratti Günther, 1892; Botia fangi Tchang, 1930] (Vermilion Botia) | `--+-- L. taeniops (Sauvage, 1878) [Cobitis Cha-ny Dabry de Thiersant, 1872; Parabotia taeniops Sauvage, 1878; Leptobotia purpurea Nichols,1925; L. taeniaps (lapsus calami)] | `-- T. tchangi Fang, 1936 `--o Botiinae |--+-- Chromobotia macracanthus (Bleeker, 1852) Kottelat, 2004 [Cobitis macracanthus Bleeker, 1852; Botia macracanthus (Bleeker, 1852)] (Clown loach; tiikerinuoliainen) | `--o Botia Gray, 1831 [Hymenphysa M'Clelland, 1839; Diacantha Swainson 1839; Hymenophysa Bleeker, 1858 (lapsus calami)] (loistonuoliaiset) | |?- B. birdi Chaudhuri, 1909 [Botia javedi Mirza & Syed, 1995] (Birdi loach; naamionuoliainen) | |?- B. dayi Hora, 1932 (Hora loach) | |?- B. lohachata Chaudhuri, 1912 (verkkonuoliainen) | |?- B. udomritthiruji Ng, 2007 [Botia sp. “Tenasserim River”] (Emperor Botia) | |-- B. rostrata Günther, 1868 [?Botia nebulosa Blyth, 1860; Botia rostrata Günther, 1868; Botia dayi Hora, 1932] (Twin-banded Loach, Ladder Loach, Dohser (India), Gangetic Loach (India), Sergeant Major Loach; touhunuoliainen) | `--+-- B. almorhae Gray, 1831 [Botia Almorhae Gray, 1831; Botia grandis Gray, 1832; Schistura maculata M'Clelland, 1839; Schistura grandis (Gray) M'Clelland, 1839; Syncrossus grandis Blyth, 1860; Botia Blythi Bleeker, 1863] (Yo-Yo Loach, Pakistani Loach, Almorha loach, Reticulate loach; mosaiikkinuoliainen) | `--+-- B. striata Narayan Rao, 1920 [Botia striata var. kolhapurensis Kalawar & Kelkar, 1958] (Striped Loach, Zebra Loach, Thin Line Loach; raitanuoliainen) | `--+-- B. dario (Hamilton, 1822) [Cobitis dario Hamilton, 1822; Gobitis geto Hamilton, 1822; Canthophrys flavicauda Swainson, 1939; Canthophrys zebra Swainson, 1939; B. macrolineata Teugels, De Vos & Snoeks, 1986] (Queen Loach, Bengal loach; vyönuoliainen) | |-- B. kubotai Kottelat, 2004 [Botia “angelicus”] (Polka Dot loach, Marble Loach, Cloud Botia, Angelicus Loach, Burmese Border Loach; harlekiininuoliainen) | `-- B. histrionica Blyth, 1860 (Burmese Loach, Golden Zebra Loach, Silver Striped Loach; klovninuoliainen) `--+--o Yasuhikotakia Nalbant, 2002 [Botia Gray, 1831 in partim] | |?- Y. longidorsalis (Taki & Doi, 1995) [Botia longidorsalis Taki & Doi, 1995] (pitkäevänuoliainen) | |?- Y. splendida (Roberts, 1995) [Botia splendida Roberts, 1995] (loistonuoliainen) | |-- Y. lecontei (Fowler, 1937) [Botia lecontei Fowler, 1937; ?Botia pulchripinnis Paysan, 1970] (Silver loach, Yellow-finned Loach; tulinuoliainen) | `--+-- Y. caudipunctata (Taki & Doi, 1995) [Botia caudipunctata Taki & Doi, 1995] (Speckletail loach; täpläpyrstönuoliainen) | `--+-- Y. modesta (Bleeker, 1865) [Botia modesta Bleeker, 1864; B. rubripinnis Sauvage, 1876] (Orangefin loach, Orange Finned Loach, Red Finned Loach, Blue Botia, Red Tailed Botia; harmaanuoliainen) | `--+-- Y. morleti (Tirant, 1885) [Botia Morleti Tirant, 1885; Botia horae Smith, 1931] (Skunk loach, Hora Loach; kaarinuoliainen) | `-- Y. eos (Taki, 1972) [Botia eos Taki, 1972] (Sun loach; aurinkonuoliainen) `--+--o Syncrossus Blyth, 1860 [Botia Gray, 1831 in partim] | |?- S. sp. (Laos Yellow Tiger Botia/Loach) | |?- S. lucasbahi (Fowler, 1937) [Botia lucas-bahi Fowler, 1937; Botia Beauforti var. formosa Pellegrin & Fang, 1940; Botia yunnanensis Chen, 1980] | |--+-- S. hymenophysa (Bleeker, 1852) [Cobitis hymenophysa Bleeker, 1852; Hymenophysa MacClellandi Bleeker, 1859; Botia hymenophysa ] (Tiger loach, Tiger Botia, Green Tiger Loach; seepranuoliainen) | | `-- S. reversus (Roberts, 1989) [Botia reversa Roberts, 1989] (pikkuseepranuoliainen) | `--+-- S. helodes (Sauvage, 1876) [Botia helodes Sauvage, 1876] (Banded Tiger loach, Tiger Botia; isoseepranuoliainen) | `--+-- S. berdmorei Blyth, 1860 [Syncrossus Berdmorei Blyth, 1860; Botia berdmorei (Blyth, 1860)] (Burmese Peppered Firetail Botia, Blyth's Loach, Tiger Botia, Berdmore's Loach; rengasnuoliainen) | `--+-- B. sp. aff. S. berdmorei | `-- S. beauforti (Smith, 1931) [Botia beauforti Smith, 1931] (Beaufort's Loach, Tiger Botia, Chameleon loach; helminuoliainen) `--+--o Ambastaia Kottelat, 2012 [Yasuhikotakia Nalbant, 2002 in partim] | |-- A. nigrolineata (Kottelat & Chu, 1987) Kottelat, 2012 [Botia nigrolineata Kottelat & Chu, 1987; Yasuhikotakia nigrolineata (Kottelat & Chu, 1987)] (Black Lined Loach; mustajuovanuoliainen) | `-- A. sidthimunki (Klausewitz, 1959) Kottelat, 2012 [Botia sidthimunki Klausewitz, 1959; Yasuhikotakia sidthimunki (Klausewitz, 1959)] (Dwarf Chain Loach, Dwarf Loach, Dwarf Botia, Chipmunk Botia, “Sid”, Ladderback loach; kääpiönuoliainen) `--o Sinibotia Fang, 1936 [Botia (Sinibotia) Fang, 1936] |?- S. longiventralis (Yang & Chen, 1992) [Botia longiventralis Yang & Chen, 1992] |?- S. reevesae (Chang, 1944) [Botia reevesae Chang, 1944] |?- S. zebra (Wu, 1939) [Botia zebra Wu, 1939] |-- S. robusta (Wu, 1939) [Botia robusta Wu, 1939; Botia hexafurca Mai, 1978] (Chinese Golden Zebra loach, Kansu loach; vankkanuoliainen) `--+-- S. superciliaris (Günther, 1892) Fang, 1936 [Botia superciliaris Günther, 1892] (Golden Chinese Loach; kultaseepranuoliainen) `-- S. pulchra (Wu, 1939) [Botia pulchra Wu, 1939; Botia gigantea Mai, 1978] (Twelve Banded Chinese Loach, Golden Chinese Loach; vannenuoliainen)
Reference(s):
- Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698 - Carroll, R. L., 1988: Appendix. 594-648
in Carroll, R. L., 1988: Vertebrate paleontology and evolution.
–W. H. Freeman and company, New York, 1988, 698 - Frickhinger, K. A., 1995: Fossil Atlas – Fishes.
–Mergus – Publishers for Natural History and Pet Books, Hans A. Baensch, Malle, Germany, 1-1088 - Froese, R. & Pauly, D. (eds), 2001: FishBase.
–iNet: www.fishbase.org - Kottelat, M., 2012: Conspectus Cobitidum: an inventory of the loaches of the world (Teleostei: Cypriniformes: Cobitoidei).
–The Raffles Bulletin of Zoology: Vol. #Supplement Number 26, pp. 1-199 - Lahti, S., Malmström, K. K., Koli, L., Leikola, A., Syrjämäki, J. & Lahti, J., 1980: Zoo, Suuri Eläinkirja 5: Kalat, sammakkoeläimet, matelijat.
–Werner Söderström Osakeyhtiö, Porvoo-Helsinki-Juva, 1980.
Original publication [in French]: Beauté Du Monde Animal: IX Reptiles/Amphibiens & X Poissons.
–Rizzoli Editore, Milano, 1968 - Nelson, J. S., 1994: Fishes of the world.
–John Wiley & Sons Inc., New York, 1994, xx-600 - Long, J. A., 1995: The Rise of Fishes: 500 million years of evolution.
–Johns Hopkins University Press, Baltimore & London, pp. 1-223 - Šlechtová, V., Bohlen, J., Freyhof, J. & Ráb, P., 2006: Molecular phylogeny of the Southeast Asian freshwater fish family Botiidae (Teleostei: Cobitoidea) and the origin of polyploidy in their evolution.
–Molecular Phylogenetics and Evolution: Vol. 39, #2, pp. 529-541 [doi: 10.1016/j.ympev.2005.09.018] - Varjo, M., 2001: Akvaario-opas. [in Finnish]
–AquaGeo & Gummerus Kirjapaino Oy, Jyväskylä, 2001, 1-143