After Carroll, 1988, Nelson, 1994, and Lovejoy & Collette, 2001, with Recent species from Froese & Pauly, 2001 [“The Fishbase”]

<== Belonidae (nokkakalat) [10 genera, 32 species]
   |?- Ablennes hians (Valenciennes, 1846) (Flat needlefish)
   |?-o Belone Plioc. Eu. Mioc. NAf. R. Oc. [2]
   |  |--o B. belone (Linnaeus, 1761) (nokkakala)
   |  |  |-- B. b. belone (Linnaeus, 1761) (Garpike)
   |  |  |-- B. b. euxini Günther, 1866
   |  |  `-- B. b. gracilis Lowe, 1839
   |  `-- B. svetovidovi Collette & Parin, 1970
   |?- Petalichthys capensis Regan, 1904 (Cape needlefish)
   |?-o Tylosurus [5]
   |  |--o T. acus
   |  |  |-- T. a. acus (Lacépède, 1803) (Agujon needlefish)
   |  |  |-- T. a. imperialis (Rafinesque, 1810)
   |  |  |-- T. a. melanotus (Bleeker, 1850) (Keel-jawed needlefish)
   |  |  |-- T. a. pacificus (Steindachner, 1876) (Pacific agujon needlefish)
   |  |  `-- T. a. rafale Collette & Parin, 1970 (Atlantic agujon needlefish)
   |  |-- T. choram (Rüppell, 1837) (Red Sea houndfish)
   |  |--o T. crocodilus
   |  |  |-- T. c. crocodilus (Péron & Lesueur, 1821) (Hound needlefish)
   |  |  `-- T. c. fodiator Jordan & Gilbert, 1882 (Mexican needlefish)
   |  |-- T. gavialoides (Castelnau, 1873)
   |  `-- T. punctulatus (Günther, 1872) (Spotted long-tom)
   `--o Strongylura [14] [paraphyletic]
      |?- Strongylura anastomella (Valenciennes, 1846)
      |?- Strongylura krefftii (Günther, 1866) (Long tom)
      |?- Strongylura leiura (Bleeker, 1850) (Banded needlefish)
      |?- Strongylura strongylura (van Hasselt, 1823) (Spottail needlefish)
      |?- Strongylura urvillii (Valenciennes, 1846)
      |--o Strongylura notata
      |  |-- S. n. forsythia Breder, 1932
      |  `-- S. n. notata (Poey, 1860) (Redfin needlefish)
      `--+-- Scomberesocidae (makrillihauet; sauries) [Nelson: 4 genera, 4 species] [Fishbase: 4/4]
         `--+--+-- Strongylura incisa (Valenciennes, 1846) (Reef needlefish)
            |  `--o Xenentodon [2]
            |     |-- X. cancila (Hamilton, 1822) (Freshwater garfish)
            |     `-- X. canciloides (Bleeker, 1853)
            `--+--o Platybelone argalus 
               |  |-- P. a. annobonensis Collette & Parin, 1970 (Annobon keeltail needlefish)
               |  |-- P. a. argalus (Lesueur, 1821) (Keeltail needlefish)
               |  |-- P. a. lovii (Günther, 1866)
               |  |-- P. a. platyura (Bennett, 1832) (Keeled needlefish)
               |  |-- P. a. pterura (Osburn & Nichols, 1916) (Baja California keeltail needlefish)
               |  `-- P. a. trachura (Valenciennes, 1846)
               `--+--+--o Pseudotylosurus [2]
                  |  |  |-- P. angusticeps (Günther, 1866)
                  |  |  `-- P. microps (Günther, 1866)
                  |  `--+-- Strongylura senegalensis (Valenciennes, 1846) (Senegal needlefish)
                  |     `-- Strongylura timucu (Walbaum, 1792) Timucu
                  `--+--o Strongylura sensu stricto?
                     |  |--+-- Strongylura fluviatilis (Regan, 1903)
                     |  |  `-- Strongylura scapularis (Jordan & Gilbert, 1882) (Shoulderspot needlefish)
                     |  `--+-- Strongylura hubbsi Collette, 1974
                     |     `--+-- Strongylura exilis (Girard, 1854) (Californian needlefish)
                     |        `-- Strongylura marina (Walbaum, 1792) (Atlantic needlefish)
                     `--+--o Belonion [2]
                        |  |-- B. apodion Collette, 1966
                        |  `-- B. dibranchodon Collette, 1966
                        `--o Potamorrhaphis [3]
                           |-- P. eigenmanni Miranda-Ribeiro, 1915
                           |-- P. guianensis (Jardine, 1843)
                           `-- P. petersi Collette, 1974

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