After Motani, 1998, 1999, Nicholls & Manabe, 2001, Chen, Motani, Cheng, Jiang & Rieppel, 2014b, and Motani, Jiang, Chen, Tintori, Rieppel, Ji & Huang, 2014
Family-level taxonomy of ichthyopterygians
<==o †Ichthyosauromorpha Motani, Jiang, Chen, Tintori, Rieppel, Ji & Huang, 2014 (ichthyosaurs and relatives; kalaliskomaiset matelijat) |?- †Wumengosaurus delictomandibularis Jiang, Rieppel, Motani, Hao, W-C. Sun, Schmitz, & Z-Y. Sun, 2008 [originally treated as nothosauroid sauropterygian] |--o †Hupehsuchia Carroll & Dong, 1991 | |-- †Nanchungosaurus suni Wang, 1959; L.Trias. SEAs. [Nanchangosauridae Wang, 1959] | `--+-- †Eohupehsuchus brevicollis Chen, Motani, Cheng, Jiang & Rieppel, 2014c | `--o †Hupehsuchidae Young, 1972 | |-- †Hupehsuchus nanchengensis Young, 1972; L.Trias. SEAs. | `--o †Parahupehsuchinae Chen, Motani, Cheng, Jiang & Rieppel, 2015 | |-- †Eretmorhipis carrolldongi Chen, Motani, Cheng, Jiang & Rieppel, 2015 [IVPP.V4070; WGSC.V26020] | `-- †Parahupehsuchus longus Chen, Motani, Cheng, Jiang & Rieppel, 2014a; L.Trias. SEAs. `--o †Ichthyosauriformes Motani, Jiang, Chen, Tintori, Rieppel, Ji & Huang, 2014 |--o †Nasorostra Jiang, Motani, Huang, Tintori, Hu, Rieppel, Fraser, Ji, Kelley, Fu & Zhang, 2016 | |-- †Cartorhynchus lenticarpus Motani, Jiang, Chen, Tintori, Rieppel, Ji & Huang, 2014 | `-- †Sclerocormus parviceps Jiang, Motani, Huang, Tintori, Hu, Rieppel, Fraser, Ji, Kelley, Fu & Zhang, 2016 `--o †Ichthyopterygia Owen, 1840 |?- †Thaisaurus chonglakmanii Mazin, Suteethorn, Buffenaut, Jaeger & Helmcke-Ingavat, 1991 |?- †Isfjordosaurus minor Motani, 1999 [Pessopteryx] |-- †Utatsusaurus hataii Shikama, Kamei & Murata, 1978 |-- †Parvinatator wapitiensis Nicholls & Brickman, 1995 `--o Eoichthyosauria Motani, 1999b [Ichthyosauria sensu lato] |--o †Grippidae Wiman, 1929 [Grippidia Wiman, 1929] | |?- †Gulosaurus helmiCuthbertson, Russell & Anderson, 2013 | |-- †Grippia longirostris Wiman, 1929 | `--o †Chaohusaurus Young & Dong, 1972 [Anhuisaurus Chen, 1985; Chensaurus (Chen, 1985) Mazin et al., 1991] | |-- †C. geishanensis Young & Dong, 1972 [Anhuisaurus faciles Chen, 1985; Chensaurus faciles (Chen, 1985) Mazin et al., 1991] | |?- †C. chaoxianensis (Chen, 1985) [Anhuisaurus chaoxianensis Chen, 1985; Chensaurus chaoxianensis (Chen, 1985) Mazin et al., 1991] | `-- †C. zhangjiawanensis Chen, Sander, Long & Wang, 2013 `-- †Ichthyosauria Blainville, 1835
Ichthyopterygia incertae sedis:
†Protoichthyosauridae Appleby, 1979 †Protoichthyosaurus prosostealis Appleby, 1979 Nomina dubia: †Svalbardosaurus crassidens Mazin, 1981 †Shastasaurus altispinus Merriam, 1902 †Shastasaurus carinthiacus Huene, 1925 †Shastasaurus careyi Merriam, 1902 †Pessopteryx arctica Wiman, 1910 †Pessopteryx nisseri Wiman, 1910 †Pessopteryx pinguis Wiman, 1910
After Jiang, Motani, Huang, Tintori, Hu, Rieppel, Fraser, Ji, Kelley, Fu & Zhang, 2016<==o †Ichthyosauromorpha |-- †Hupehsuchia `--o †Ichthyosauriformes |--o †Nasorostra | |-- †Cartorhynchus | `-- †Sclerocormus `--o †Ichthyopterygia |-- †Chaohusaurus geishanensis |-- †Chaohusaurus chaoxianensis |-- †Chaohusaurus zhangjiawanensis `--+-- †Utatsusaurus |-- †Parvinatator |--o †Grippidia | |-- †Grippia | `-- †Gulosaurus `-- †Ichthyosauria
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After Motani, Jiang, Chen, Tintori, Rieppel, Ji & Huang, 2014 (C. Ji et al., in prep.)<==o †Ichthyosauromorpha |-- †Hupehsuchia `--o †Ichthyosauriformes |-- †Cartorhynchus `--o †Ichthyopterygia |--o †Chaohusaurus | |-- †C. geishanensis | |-- †C. chaoxianensis | `-- †C. zhangjiawanensis `--+--o †Grippidia | |-- †Utatsusaurus | |-- †Parvinatator | `--+-- †Grippia | `-- †Gulosaurus `-- †Ichthyosauria
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After Chen, Motani, Cheng, Jiang & Rieppel, 2014b<==o |-- †Thalattosauriformes `--+-- †Wumengosaurus `--+-- †Ichthyopterygia `--+-- †Nanchungosaurus `-- †Hupehsuchus
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Reference(s):
- Bardet, N. & Fernandez, M., 2000: A new ichthyosaur from the Upper Jurassic lithographic limestones of Bavaria.
–Journal of Paleontology: Vol. 74, #3, pp. 503-511 - Chen, X.-h., Motani, R., Cheng, L., Jiang, D.-y. & Rieppel, O., 2015: A New Specimen of Carroll's Mystery Hupehsuchian from the Lower Triassic of China.
–Plos One: Vol. 10, #5, pp. e0126024 (1-15) [doi: 10.1371/journal.pone.0126024] - Chen, X.-h., Motani, R., Cheng, L., Jiang, D.-y. & Rieppel, O., 2014a: A Carapace-Like Bony ‘Body Tube’ in an Early Triassic Marine Reptile and the Onset of Marine Tetrapod Predation.
–Plos One: Vol. 9, #4, pp. e94396 (1-8) [doi: 10.1371/journal.pone.0094396] - Chen, X.-h., Motani, R., Cheng, L., Jiang, D.-y. & Rieppel, O., 2014b: The Enigmatic Marine Reptile Nanchangosaurus from the Lower Triassic of Hubei, China and the Phylogenetic Affinities of Hupehsuchia.
–Plos One: Vol. 9, #7, pp. e102361 (1-13) [doi: 10.1371/journal.pone.0102361] - Chen, X.-h., Motani, R., Cheng, L., Jiang, D.-y. & Rieppel, O., 2014c: A Small Short-Necked Hupehsuchian from the Lower Triassic of Hubei Province, China.
–Plos One: Vol. 9, #12, pp. e115244 [doi: 10.1371/journal.pone.0115244] - Chen, X., Sander, P. M., Long, C. & Wang, X., 2013: A new Triassic primitive ichthyosaur from Yuanan, South China.
–Acta Geologica Sinica - English Edition: Vol. 87, #3, pp. 672-677 [doi: 10.1111/1755-6724.12078] - Cuthbertson, R. S., Russell, A. P. & Anderson, J. S., 2013: Cranial morphology and relationships of a new grippidian (Ichthyopterygia) from the Vega-Phroso Siltstone Member (Lower Triassic) of British Columbia, Canada.
–Journal of Vertebrate Paleontology: Vol. 33, #4, pp. 831-847 [doi: 10.1080/02724634.2013.755989] - Fernández, M. S., 1999: a new ichthyosaur from the Los Molles Formation (Early Bajocian), Neuquen Basin, Argentina.
–Journal of Paleontology: Vol. 73, #4, pp. 677-681 - Jiang, D.-Y., Motani, R., Huang, J.-D., Tintori, A., Hu, Y.-C., Rieppel, O., Fraser, N. C., Ji, C., Kelley, N. P., Fu, W.-L. & Zhang, R., 2016: A large aberrant stem ichthyosauriform indicating early rise and demise of ichthyosauromorphs in the wake of the end-Permian extinction.
–Scientific Reports: Vol. 6, Art. nr. 26232 [doi: 10.1038/srep26232] - Jiang, D.-Y., Rieppel, O., Motani, R., Hao, W.-C., Sun, Y.-L., Schmitz, L. & Sun, Z.-Y., 2008: A New Middle Triassic Eosauropterygian (Reptilia, Sauropterygia) from Southwestern China.
–Journal of Vertebrate Paleontology: Vol. 28, #4, pp. 1055-1062 [doi: 10.1671/0272-4634-28.4.1055] - McGowan, C., 1994: A new species of Shastasaurus (Reptilia: Ichthyosauria) from the Triassic of British Colombia: The most complete exemplar of the genus.
–Journal of Vertebrate Paleontology: Vol. 14, #2, pp. 168-179 - McGowan, C., 1997: The taxonomic status of the Late Jurassic ichthyosaur Grendelius mordax: A preliminary report.
–Journal of Vertebrate Paleontology: Vol. 17, #2, pp. 428-430 - McGowan, C. & Milner, A. C., 1999: A new Pliensbachian ichthyosaur from Dorset, England.
–Palaeontology: Vol. 42, #5, pp. 761-768 - Motani, R., 1996: Redescription of the dental features of an Early Triassic ichthyosaur, Utatsusaurus hataii.
–Journal of Vertebrate Paleontology: Vol. 16, #3, pp. 396-402 - Motani, R., 1997: Redescription of the dentition of Grippia longirostris (Ichthyosauria) with a comparison with Utatsusaurus hataii.
–Journal of Vertebrate Paleontology: Vol. 17, #1, pp. 39-44 - Motani, R., 1999: The skull and taxonomy of Mixosaurus (Ichthyopterygia).
–Journal of Paleontology: Vol. 73, #5, pp. 924-935 - Motani, R., 1999: On the evolution and homologies of ichthyopterygian forefins.
–Journal Of Vertebrate Paleontology: Vol. 19, #1, pp. 28-41. - Motani, R., 1999: Phylogeny of the Ichthyopterygia.
–Journal Of Vertebrate Paleontology: Vol. 19, #3, pp. 473-496. - Motani, R., 2000: Skull of Grippia longirostris: No contradiction with a diapsid affinity for the Ichthyosauria.
–Palaeontology: Vol. 43, #1, pp. 1-14 - Motani, R., 2000: Is Omphalosaurus ichthyopterygian? - A phylogenetic perspective.
–Journal Of Vertebrate Paleontology: Vol. 20, #2, pp. 295-301 - Motani, R., Jiang, D.-Y., Chen, G.-B., Tintori, A., Rieppel, O., Ji, C. & Huang, J.-D., 2014: A basal ichthyosauriform with a short snout from the Lower Triassic of China.
–Nature: in press [doi: 10.1038/nature13866] - Motani, R., Jiang, D.-Y., Tintori, A., Rieppel, O., Chen, G.-B. & You, H., 2015: Status of Chaohusaurus chaoxianensis (Chen, 1985).
–Journal of Vertebrate Paleontology: Vol. 35, #1, pp. e892011 (1-8) [doi: 10.1080/02724634.2014.892011] - Motani, R., Minoura, N. & Ando, T., 1998: Ichthyosaurian relationships illuminated by new primitive skeletons from Japan.
–Nature: Vol. 393, 21 May. pp. 255-257 - Motani, R. & You, H., 1998: Taxonomy and limb ontogeny of Chaohusaurus geishanensis (Ichthyosauria), with a note on the allometric equation.
–Journal of Vertebrate Paleontology: Vol. 18, #3, pp. 533-540 - Motani, R., You, H., & McGowan, C., 1996: Eel-like swimming in the earliest ichthyosaurs.
–Nature: Vol. 382, 25 July, pp. 347-348 - Nicholls, E. L. & Manabe, M., 2001: A new genus of ichthyosaur from the Late Triassic Pardonet Formation of British Colombia: bridging the Triassic Jurassic gap.
–Canadian Journal of Earth Sciencies: Vol. 38, #6, pp. 983-1002 - Sander, P. M., 1989: The large ichthyosaur Cymbospondylus buchseri, sp. nov., from the Middle Triassic of Monte Giorgio (Switzerland), with a survay of the genus in Europe.
–Journal of Vertebrate Paleontology: Vol. 9, #2, pp. 163-173 - Storrs, G. W., Arkhangel'skii, M. S. & Efimov, V. R., 2000: Mezosoic marine reptiles of Russia and other former Soviet republics. 187-210
in Benton, M. J., Shishkin, M. A., Unvin, D. M. & Kurochkin, E. N., (eds.) 2000: The Age of Dinosaurs in Russia and Mongolia.
–Cambridge University Press, Cambridge, 2000 xxxix-696