After Meylan & Ganko, 1997, Lyson & Joyce, 2010, and Lyson & Joyce, 2011
<==o †Paracryptodira [=? Pleurosternoidea Hay, 1930] |?-o †Solemydidae de Lapparent de Broin & Murelaga, 1996 | |?- †Plastremys lata Owen, in Parkinson, 1881 [Hylaeochelys lata (Owen) Milner, 2004] | |--o †Helochelydra Nopcsa, 1928 [=? †Helochelys] | | |-- †H. nopcsai de Lapparent de Broin & Murelaga, 1999 | | |-- †H. bakewelli (Mantell, 1833) Milner, 2004 | | `-- †H. anglica (Lydekker, 1889b) Milner, 2004 | |-- †Naomichelys speciosa Hay, 1908 | `-- †Solemys vermiculata de Lapparent de Broin and Murelaga 1996 |--o †Pleurosternidae Cope, 1868 [incl. ?Chengyuchelyidae Yeh, 1990] | |?-o †Chengyuchelys Young & Chow, 1953 [Chengyuchelyidae Yeh, 1990] | | |-- †C. baenoides Young & Chow, 1953 | | `-- †C. zigongensis Yeh 1982 | |-- †Dorsetochelys delairi Evans & Kemp, 1976 [Cryptodira incertae sedis (Milner, 2004)] | |?- †Uluops uluops Bakker, 1990 | |--o †Pleurosternon Owen, 1853 [taxonomic history] | | |-- †P. bullockii (Owen, 1842) Lydekker & Boulenger, 1887 [taxonomic history] | | `-- †P. portlandicum Lydekker, 1889 | |?-o †Compsemys Leidy, 1856; E.Cret.-Pal. NAm. Eu. | | |-- †C. bakewelli (Mantell, 1833) [Helochelydra bakewelli; Tretosternum bakewelli; Trionyx bakewelli] | | `-- †C. victa Leidy, 1856 [Compsemys obscura Leidy, 1856; Compsemys parva Hay, 1910; Compsemys puercensis Gilmore, 1919; Compsemys torrejonensis Gilmore, 1919; Compsemys vafer Hay, 1910; Emys obscurus Leidy, 1856; Glyptops depressus Hay, 1908] | `--+--o †“Glyptopsinae” Marsh, 1890 [Glyptopsidae Marsh, 1890] | | |?- †Mesochelys durlstonensis Evans & Kemp, 1975 [treated as a synonym of Pleurosternon by Milner (2004)] | | `--o †Glyptops Marsh, 1890 [taxonomic history] | | |?- †“Glyptops” typocardium (Seeley, 1869) Milner, 2004 [taxonomic history] | | |-- †G. plicatulus (Cope, 1877) Marsh, 1890 [taxonomic history] | | `-- †G. ruetimeyeri (Lydekker, 1889b) [taxonomic history] | `--o †“Dinochelinae” | |-- †Desmemys bertelsmanni Wegner, 1911 | `-- †Dinochelys whitei Gaffney, 1979 `--o †Baenoidea [Baenidae sensu lato] [paraphyletic??] |?- †Macrobaenidae Sukhanov, 1964 `--o †Baenidae Cope, 1882 sensu stricto |?- †Goleremys mckennai Hutchison, 2005 |-- †Arundelemys dardeni Lipka, Therrien, Weishampel, Jamniczky, Joyce, Colbert & Brinkman, 2006 `--+-- †Trinitichelys hiatti Gaffney, 1972 `--+--o †Neurankylus Lambe, 1902 [Charitemys Hay, 1908] [Neurankylidae] | |-- †N. eximius Lambe, 1902 [Baena fluviatilis Parks 1933; Charitemys captans Hay, 1908; Neurankylus baueri Gilmore, 1916; Thescelus fluviatilis Parks, 1933] | `-- †N. wyomingensis Gilmore, 1919 `--+-- †Hayemys latifrons Gaffney, 1972 |-- †Thescelus insiliens Hay, 1908 [Baena longicauda Russell, 1934; Thescelus hemispherica Gilmore, 1935; Thescelus rapiens Hay, 1908] |-- †“Denazinemys” ornata (Gilmore, 1935) Lucas & Sullivan, 2006 [Baena ornata Gilmore, 1935] `--o †Baenodda Gaffney & Meylan, 1988 [Eubaeninae] |?- †“Baena” hatcheri Hay, 1901 |?- †“Baena” marshi Hay, 1904 |--+-- †“Baena” hayi Gilmore, 1916 | `--+--o †Plesiobaena Gaffney, 1972 | | |-- †P. antiqua (Lambe, 1902) [Baena antiqua Lambe, 1902; Baena antiqua Lambe; Russell, 1934; Plesiobaena antigua (Lambe); Gaffney, 1972 (in partim)]; U. Cret. NA. | | `--+-- †P. sp. U. Cret. NA. [Maastrichian species (Brinkman, 2003)] | | `-- †P. putorius Gaffney, 1972; Pal. NA. [=? Cedrobaena putorius] | `--+-- †Peckemys brickman | `--+-- †Cedrobaena putorius [=? Plesiobaena putorius Gaffney, 1972] | `--+-- †Gamerabaena sonsalla Lyson & Joyce, 2010 | `--o †Palatobaena | |-- †P. cohen Lyson & Joyce, 2009 | `--+-- †P. gaffneyi | `-- †P. bairdi `--+--+-- †Stygiochelys estesi Gaffney & Hiatt, 1971 | `--+-- †Baena arenosa Leidy, 1890 | `--o †Chisternon Leidy, 1872 | |-- †C. undatum (Leidy, 1871) | `-- †C. hebraicum Cope 1872 `--+-- †Eubaena cephalica (Hay, 1904) Hay, 1908 [Baena cephalica Hay, 1904] `--+-- †Denazinemys nodosa (Gilmore, 1916) Lucas & Sullivan, 2006 [Baena nodosa Gilmore, 1916] `--o †Boremys Lambe, 1906 |-- †B. pulchra Lambe, 1906 `-- †B. grandis Gilmore, 1935
After Lyson & Joyce, 2011<==o †Paracryptodira |-- †Dinochelys whitei `--+-- †Compsemys victa `--+-- †Uluops uluops |--o †Pleurosternidae | |-- †Dorsetochelys delairi | `--+-- †Pleurosternon | `-- †Glyptops `--o †Baenidae |-- †Arundelemys dardeni `--+-- †Trinitichelys hiatti `--+-- †Neurankylus eximius `--+-- †Hayemys latifrons |-- †“Denazinemys” ornata |--+--+-- †Stygiochelys estesi | | `--+-- †Baena arenosa | | `-- †Chisternon undatum | `--+-- †Eubaena cephalica | `--+-- †Denazinemys nodosa | `--o †Boremys | |-- †B. pulchra | `-- †B. grandis `--+-- †Plesiobaena antiqua `--+-- †Peckemys brickmani `--+-- †Cedrobaena putorius `--+-- †Gamerabaena sonsalla `--o †Palatobaena |-- †P. cohen `--+-- †P. gaffneyi `-- †P. bairdi Reference(s):
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After Lyson & Joyce, 2010<==o †Paracryptodira |-- †Pleurosternon |-- †Glyptops `--o †Baenidae |--+-- †Trinitichelys hiatti | `-- †Neurankylus eximius `--+-- †Thescelus insiliens |-- †Hayemys latifrons `--o †Baenodda |--+-- †“Baena” hayi | `--+-- †Plesiobaena antiqua | `--+-- †Gamerabaena sonsalla | `--o †Palatobaena | |-- †P. cohen | `--+-- †P. gaffneyi | `-- †P. bairdi `--+--+-- †Boremys pulchra | |-- †Boremys grandis | |-- †Eubaena cephalica | `-- †“Baena” hatcheri `--+-- †Stygiochelys estesi `--+-- †Baena arenosa `-- †Chisternon undatum Reference(s):
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After Lipka, Therrien, Weishampel, Jamniczky, Joyce, Colbert & Brinkman, 2006<==o †Paracryptodira |-- †Arundelemys dardeni `--+--o †Pleurosternidae | |-- †Pleurosternon | `--+-- †Glyptops | `-- †Dinochelys whitei `--o †Baenidae |-- †Neurankylus eximius `--+-- †Trinitichelys hiatti `--+-- †Plesiobaena antiqua `--+-- †Boremys pulchra `--+-- †Baena arenosa `-- †Chisternon undatum Reference(s):
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Reference(s):
- Brinkman, D. B., 2003: Anatomy and systematics of Plesiobaena antiqua (Testudines: Baenidae) from the Mid-Campanian Judith River Group of Alberta, Canada.
–Journal of Vertebrate Paleontology: Vol. 23, #1, pp. 146-155 - Brickman, D. B., & Nicholls, E. L., 1993: The skull of Neurankylus eximius (Testudines: Baenidae) and a reinterpretation of the relationships of this taxon.
–Journal of Vertebrate Paleontology: Vol. 13, #3, pp. 273-281 - Brickman, D. B., Stadtman, K. & Smith, D., 2000: New material of Dinochelys whitei Gaffney, 1979, from the Dry Mesa Quarry (Morrison Formation, Jurassic) of Colorado.
–Journal of Vertebrate Paleontology: Vol. 20, #2, pp. 269-274 - Lipka, T. R., Therrien, F., Weishampel, D. B., Jamniczky, H. A., Joyce, W. G., Colbert, M. W. & Brinkman, D. B., 2006: A new turtle from the Arundel clay facies (Potomac Formation, Early Cretaceous) of Maryland, U.S.A.
–Journal of Vertebrate Paleontology: Vol. 26, #2, pp. 300-307 [doi: 10.1671/0272-4634(2006)26[300:antfta]2.0.co;2] - Lyson, T. R. & Joyce, W. G., 2010: A new baenid turtle from the upper Cretaceous (Maastrichtian) Hell Creek Formation of North Dakota and a preliminary taxonomic review of Cretaceous Baenidae.
–Journal of Vertebrate Paleontology: Vol. 30, #2, pp. (394-402) [doi: 10.1080/02724631003618389] - Lyson, T. R. & Joyce, W. G., 2011: Cranial Anatomy and Phylogenetic Placement of the Enigmatic Turtle Compsemys victa Leidy, 1856.
–Journal of Paleontology: Vol. 85, #4, pp. 789-801 [doi: 10.1666/10-081.1] - Meylan, P. A. & Ganko, E., 1997: Higher Relationships of Testunides – The extinct and living turtles.
–iNet: The Tree of Life: [no longer available] - Milner, A. R., 2004: The turtles of the Purbeck Limestone Group of Dorset, southern England.
–Palaeontology: Vol. 47, #6, pp. 1441-1467 - Sukhanov, V. R., 2000: Mesozoic turtles of Middle and Central Asia. 309-367
in Benton, M. J., Shishkin, M. A., Unvin, D. M. & Kurochkin, E. N., (eds.) 2000: The Age of Dinosaurs in Russia and Mongolia.
–Cambridge University Press, Cambridge, 2000, xxxix-696 - Zonneveld, J.-P., Gunnell, G. F. & Bartels, W. S., 2000: Early Eocene fossil vertebrates from the southwestern Green River basin, Lincoln and Uinta Counties, Wyoming.
–Journal of Vertebrate Paleontology: Vol. 20, #2, pp. 369-386