Mikael Fortelius, FT



Evoluutiopaleontologian professori
–paleontologian ja paleoekologian linja

Kristine Bonnevie Professor, Centre for Ecological and Evolutionary Synthesis (CEES), University of Oslo

Geotieteiden ja maantieteen laitos
PL 64 (Gustaf Hällströmin katu 2a)
00014 Helsingin yliopisto

puh: 02941 50837
sposti:  mikael.fortelius(at)helsinki.fi

Pääasiallinen opetusalue

  • varhaisihmisen elinympäristöjen olosuhteet ja nisäkäsyhteisöt
  • historiallinen geologia
  • yleinen paleontologia
  • muinaiset ympäristöt ja ekosysteemit
  • toiminnallinen morfologia
  • nisäkkäiden ja ihmisen evoluutio


  • Euraasian maanisäkkäiden ja mannerympäristöjen kehitys neogeeniajalla
  • Hampaiden kehitysbiologia, toiminta ja evoluutio
  • kasvinsyöjänisäkkäiden ekomorfologia
  • nisäkäshampaan yksilönkehitys, toiminta ja evoluutio
  • skaalausongelmat

Viimeaikaiset julkaisut

Raia, P., Passaro, F., Carotenuto, F., Maiorino, L., Piras, P., Teresi, L., Meirin, S., Itescu, Y., Novosolov, M., Baiano, R.M., Fortelius, M. 2015. Cope’s Rule and the universal scaling law of ornament complexity. The American Naturalist 186, electronic publication ahead of print.

Fortelius M., Geritz S., Gyllenberg M., Toivonen, J. 2015. Adaptive dynamics on an environmental gradient that changes over a geological time-scale. Journal of Theoretical Biology 376: 91-104.

Sukselainen, L., Fortelius, M., Harrison, T. 2015. Co-occurrence of pliopithecoid and hominoid primates in the fossil record: An ecometric analysis. Journal of Human Evolution xxx:1-17. Online publication: 14-MAY-2015, DOI: 10.1016/j.jhevol.2015.04.009

Kaakinen A., Abdul Aziz, H., Passey B.H., Zhang Z., Liu L., Salminen J., Wang, L., Krijgsman W., Fortelius M. 2015. Age and stratigraphic context of Pliopithecus and associated fauna from Miocene sedimentary strata at Damiao, Inner Mongolia, China. Journal of Asian Earth Sciences 100 (2015) 78–90.

Saarinen, J.J., Boyer, A.G., Brown, J.H, Costa, D.P., Ernest, S.K.M., Evans, A.R., Fortelius, M., Gittleman, J.L., Hamilton, M.J., Harding, L.E., Lintulaakso, K., Lyons, S.K., Okie, J.G., Sibly, R.M., Stephens, P.R., Theodor, J., Uhen, M.D., Smith, F.A. 2014. Patterns of maximum body size evolution in Cenozoic land mammals: eco-evolutionary processes and abiotic forcing. Proceedings of the Royal Society B 281, doi: 10.1098/rspb.2013.2049.

Müller J., Clauss M., Codron D., Schulz E., Hummel J., Fortelius M., Kircher P., Hatt J.-M. 2014. Growth and wear of incisor and cheek teeth in domestic rabbits (Oryctolagus cuniculus) fed diets of different abrasiveness. Journal of Experimental Zoology A 321: 283-298.

Barnosky, A.D., Hadly, E. A., Dirzo, R., Fortelius, M. & Stenseth, N.C. 2014.  Translating science for decision makers to help navigate the Anthropocene.  The Anthropocene Review 1:160–170. doi: 10.1177/2053019614533670

Fortelius, M., Eronen, J.T., Kaya, F., Tang, H., Raia, P. & Puolamäki, K. 2014. Evolution of Neogene mammals in Eurasia: environmental forcing and biotic interactions. Annual Reviews of Earth and Planetary Science 42:579–604, doi: 10.1146/annurev-earth-050212-124030.

Barnosky A.D., Brown J.H., Daily G.C., et al. 2014. Introducing the Scientific Consensus on Maintaining Humanity's Life Support Systems in the 21st Century: Information for Policy Makers. The Anthropocene Review 1:78-109.

Cardoso, P.M., Rigal, F., Carvalho, J.C., Fortelius, M., Borges, P.A.V., Podani, J., Schmera, D. 2014. Partitioning taxon, phylogenetic and functional beta diversity into replacement and richness difference components. Journal of Biogeography 41:749–761.

Wang, X., L.F.  Flynn and M. Fortelius (Eds). 2013. Fossil Mammals of Asia. Neogene terrestrial biostratigraphy and chronology. Columbia University Press. 732 pp.

Raia, P. & Fortelius, M. 2013.  Cope’s Law of the Unspecialized, Cope’s Rule, and weak directionality in evolution. Evolutionary Ecology Research 15: 747–756. 

Nummela S., Pihlström, H., Puolamäki, K., Fortelius, M., Hemilä, S., Reuter, T. 2013. Exploring the mammalian sensory space: co-operations and trade-offs among senses. Journal of Comparative Physiology. A, Neuroethology, Sensory, Neural, and Behavioral Physiology 199:1077-1092.

Okie J.G., Boyer A.G., Brown J.H., Costa D.P., Ernest S.K.M., Evans A.R., Fortelius M., Gittleman J.L., Hamilton M.J., Harding L.E., Lintulaakso K., Lyons S.K., Saarinen J.J., Smith F.A., Stephens P.R., Theodor J., Uhen M.D., Sibly R.M. 2013. Effects of allometry, productivity and lifestyle on rates and limits of body size evolution. Proceedings of the Royal Society B 280: 20131007 (online June 13).

Zhang, Z., Kaakinen, A. P., Liu, L., Lunkka, J.P., Sen, S., Gose, W.A., Qiu, Z. Zheng, S.  & Fortelius, M. 2013. Mammalian biochronology of the Late Miocene Bahe Formation. Pp. 187-202 in Wang, X., Flynn, L.J.  & Fortelius, M (Eds), Fossil Mammals of Asia, New York: Columbia University Press.

Kaakinen, A. P., Passey, B. H., Zhang, Z., Pesonen, L., Liu, L. & Fortelius, M. 2013 Stratigraphy and paleoecology of the classical dragon bone localities of Baode County, Shanxi Province, China. Pp. 203-217 in Wang, X., Flynn, L.J.  & Fortelius, M (Eds), Fossil Mammals of Asia, New York: Columbia University Press.

Mirzaie Ataabadi, M., Liu, L., Eronen, J. T., Bernor, R. & Fortelius, M. 2013. Continental Scale Patterns in Neogene Mammal Community Evolution and Biogeography: A Europe-Asia Perspective. Pp. 629-655 in Wang, X., Flynn, L.J.  & Fortelius, M (Eds), Fossil Mammals of Asia, New York: Columbia University Press.

Hadly E.A., Barnosky A.D., Fortelius M., Stenseth N.C. 2013. Getting the word out on biosphere crisis. Nature 497:565. 

Fortelius, M. 2013. Commentary: The grassiness of all flesh. Journal of Biogeography 40: 1213–1214.

Uhen, M. D., Barnosky, A. D., Bills, B., Blois, J., Carrano, M. T., Carrasco, M. A., Erickson, G. M., Eronen, J. T., Fortelius, M., Graham, R. W., Grimm, E. C., O'Leary, M. A., Mast, A., Piel, W. H., Polly, P. D. & Säilä, L. K. 2013. From card catalogs to computers: Databases in vertebrate paleontology. Journal of Vertebrate Paleontology. 33:13-28.

Raia P, Carotenuto F, Passaro F, Piras P, Fulgione D, Werdelin L, Saarinen J, Fortelius M. 2013. Rapid action in the Palaeogene: the relationship between phenotypic and taxonomic diversification in Coenozoic mammals. Proceedings of the Royal Society B 280: 2012-2244.

Kaiser, T.M., Müller, D.W.H., Fortelius, M., Schultz, E., Codron, D. and Clauss, M. 2013. Hypsodonty and tooth facet development in relation to diet and habitat in herbivorous ungulates: implications for understanding tooth wear. Mammal Review 43:34–46.

Tang, H., Eronen, J. T., Micheels , A., Ahrens, B. & Fortelius, M. 2012. Asynchronous responses of East Asian and Indian summer monsoons to mountain uplift shown by regional climate modelling experiments. Climate Dynamics 40:1531-1549.

Eronen, J.T., Fortelius, M., Micheels, A., Portmann, F.T., Puolamäki, K. & Janis, C.M. 2012. Neogene aridification of the Northern Hemisphere. Geology 40:823-826.

Liu, L., Puolamäki, K., Eronen, J. T., Mirzaie Ataabadi, M., Hernesniemi, E. & Fortelius, M. 2012 Dental functional traits of mammals resolve productivity in terrestrial ecosystems past and present. Proceedings of the Royal Society B 279:2793-2799.

Wilson, G. P., Evans, A. R., Corfe, I. J., Smits, P. D., Fortelius, M. & Jernvall, J. 2012. Adaptive radiation of multituberculate mammals before the extinction of dinosaurs. Nature 483:457-460.

Heikinheimo, H., Eronen, J. T., Sennikov, A., Preston, C., Oikarinen, E., Uotila, P., Mannila, H. & Fortelius, M. 2012. Convergence in the distribution patterns of Europe's plants and mammals is due to environmental forcing. Journal of Biogeography 39:1633-1644.

Barnosky, A.D., Hadly, E.A., Bascompte, J., Berlow, E. L., Brown, J.H., Fortelius, M., Getz, W.M., Harte, J., Hastings, A., Marquet, P.A., Martinez, N.D., Mooers, A., Roopnarine, P., Vermeij, G., Williams, J.W., Gillespie, R., Kitzes, J., Marshall, C., Matzke, N., Mindell, D.P., Revilla, E. & Smith, A.B. 2012. Approaching a state shift in Earth’s biosphere. Nature 486:52-58.

Evans, A.R., Jones, D., Boyer, A.G., Brown, J.H., Costa, D.P., Dayan, T., Ernest, S.K.M.,  Fitzgerald, E.M.G. Fortelius, M.  Gittleman, J.L. Hamilton, M.J., Harding, L.E.Lintulaakso, K., Lyons, S.K., Okie, J.G. Saarinen, J.J., Sibly, R.M.,  Smith, F.A., Stephens, P.R., Theodor, J., and Uhen, M.D. 2012. Maximum rates of body size macroevolution in mammals. PNAS 109:4187-4190.

Raia P., Carotenuto F., Passaro F., Fulgione D. and Fortelius, M. 2012. Ecological specialization in fossil mammals explains Cope's rule. American Naturalist 179:328-337.

Eronen, J.T., Puolamäki, K., Heikinheimo, H., Lokki, H., Venäläinen, A., Mannila, H. and Fortelius, M. 2011. The effect of scale, climate and environment on species richness and spatial distribution of Finnish birds. Annales Zoologici Fennici 48:257-274.

Eronen, J. T., Evans, A. R., Fortelius, M. & Jernvall, J. 2011. Genera are often better than species for detecting evolutionary change in the fossil record: A reply to Salesa et al. Evolution 65:1514-1516.

Zhang, Z.Q., Wang, L.H., Kaakinen, A., Liu, L.P. and Fortelius, M. 2011. Miocene mammalian faunal succession from Damiao, central Nei Mongol and the environmental changes. Quaternary Sciences 31:608-613.

Deng, T., X. Wang, M. Fortelius, Q. Li, Y. Wang, Z.J. Tseng, G.T. Takeuchi, J. E. Saylor, L.K. Säilä, G. Xie. 2011. Out of Tibet: Pliocene woolly rhino suggests high-plateau origin of Ice Age megaherbivores. Science 333:1285-1288.

Flynn, L.F., X. Wang and M. Fortelius. 2011. Neogene terrestrial mammalian biostratigraphy and chronology in Asia: a special issue [Preface]. Vertebrata PalAsiatica 49 (3):253-256.

Wang, X. G. Xie, Z. Qiu,  Q. Li Z.J. Tseng, G.T. Takeuchi B. Wang,  M. Fortelius, A. Rosenström-Fortelius, H.  Wahlquist. Early explorations of Qaidam Basin (Tibetan Plateau) by Birger Bohlin - reconciling classic vertebrate fossil localities with modern biostratigraphy. Vertebrata PalAsiatica 49:285-310.

Tang, H., Eronen, J. T., Micheels, A. & Fortelius, M. 2011.  A regional climate model experiment to investigate the Asian monsoon in the Late Miocene. Climate of the Past 7:847-868.

Micheels, A., Bruch, A., Eronen, J. T., Fortelius, M., Harzhauser, M., Utescher, T. & Mosbrugger, V. 2011. Analysis of heat transport mechanisms from a Late Miocene model experiment with a fully-coupled atmosphere–ocean general circulation model. Palaeogeography, Palaeoclimatology, Palaeoecology 304:337-350.

Raia, P., F. Carotenuto, J. T. Eronen and M. Fortelius. 2011. Longer in the tooth, shorter in the record? The evolutionary correlates of hypsodonty in Neogene ruminants. Proceedings of the Royal Society B 278, p. 3474-3481.

Kallio, A., Puolamäki, K., Fortelius, M., and Mannila, H., 2011. Correlations and co-occurrences of taxa: the role of temporal, geographic, and taxonomic restrictions. Palaeontologia Electronica Vol. 14, Issue 1; 4A:14p;

Hernesniemi, E., I.X. Giaourtsakis, A. Evans and M. Fortelius. 2011. Rhinocerotidae. Pp. 275-294 in: T. Harrison (Ed), Paleontology and geology of Laetoli: human evolution in context. Vol. 2: Fossil hominins and the associated fauna. Springer.

Hernesniemi, E., Blomstedt, K. & Fortelius, M. 2011. Multi-view stereo 3D reconstruction of lower molars of Recent and Pleistocene rhinoceroses for mesowear analysis. Palaeontologia Electronica 14, 2, 15 pp.

Wang, X., Xie, Z., Qiu, Z., Li, Q., Tseng, Z. J., Takeuchi, G. T., Wang, B., Fortelius, M., Rosenström-Fortelius, A. & Wahlquist, H. 2011. Early explorations of Qaidam Basin (Tibetan Plateau) by Birger Bohlin - reconciling classic vertebrate fossil localities with modern biostratigraphy. Vertebrata PalAsiatica 49:285-310.

Mirzaie Ataabadi, M., Mohammadalizadeh, J., Zhang, Z., Watabe, M., Kaakinen, A. P. & Fortelius, M. 2011. Late Miocene large mammals from Ivand (Northwestern Iran). Geodiversitas. 33:709-728.

Polly, P.D., J.T. Eronen, M. Fred, G.P. Dietl, V. Mosbrugger, C. Scheidegger, D.C. Frank, J. Damuth, N.C. Stenseth, and M. Fortelius.  2011. History Matters: ecometrics and integrative climate change biology.  Proceedings of the Royal Society B, 278: 1131-1140.

Saarinen, J., E. Oikarinen, M. Fortelius, and H. Mannila. 2010. The living and the fossilised  –  how well do unevenly distributed points capture the faunal information in a grid? Evolutionary Ecology Research 12: 363–376.

Eronen, J.T., Puolamäki, K., Liu, L., Lintulaakso, K., Damuth, J., Janis, C., and Fortelius, M. 2010. Precipitation and large herbivorous mammals, part I: Estimates from present-day communities. Evolutionary Ecology Research 12, 217-233.

Eronen, J.T., Puolamäki, K., Liu, L., Lintulaakso, K., Damuth, J., Janis, C., and Fortelius, M. 2010. Precipitation and large herbivorous mammals , part II: Application to fossil data. Evolutionary Ecology Research 12, 235-248.

Eronen, J.T. , Evans, A.R., Jernvall, J., Fortelius, M. 2010. The impact of regional climate on the evolution of mammals. A case study using fossil horses. Evolution 64, 398-408. doi:10.1111/j.1558-5646.2009.00830.x

Eronen, J.T., Polly, P.D., Fred, M., Damuth, J., Frank, D.C., Mosbrugger, V., Scheidegger, C., Stenseth, N.C., Fortelius, M. 2010. Ecometrics: The traits that bind the past and present together. Integrative Zoology 5: 88-101.

Smith, F. A., Boyer, A. G., Brown, J. H., Costa, D. P., Dayan, T., Ernest, S. K. M., Evans, A. R., Fortelius, M., Gittleman, J. L., Hamilton, M. J., Harding, L. E.,Lintulaakso, K., Lyons, S. K., McCain, C., Okie, J. G., Saarinen, J. J., Sibly, R. M., Stephens, P. R., Theodor, J. & Uhen, M. D. 2010. The evolution of maximum body size of terrestrial mammals. Science. 330:1216-1219

Eronen, J. T.,  M. Mirzaie Ataabadi, A. Micheels, A. Karme, R. L. Bernor, and M. Fortelius. 2009. Distribution history and climatic controls of the Late Miocene Pikermian chronofauna. PNAS 106 (29):11867–11871.

Liu, L., J. T. Eronen, M. Fortelius. 2009.  Significant mid-latitude aridity in the middle Miocene of East Asia. Palaeogeography, Palaeoclimatology, Palaeoecology 279:201-206.

Liow, L. H., M. Fortelius; K. Lintulaakso; H. Mannila & N. C. Stenseth. 2009. Lower extinction risk in sleep-or-hide mammals. The American Naturalist 178:264-272.

Valikoitu julkaisuluettelo (pdf)

Curriculum Vitae

CV (pdf)